814651160c815d6f30776cbafe9fd268 pmc_31784545.pdf 79c70aa313d9309b6371d7fe36496dac1f6d9b59 pmc_31784545.pdf 1d7205d86b4d77c85e5f0cf8bd0b315205a0afcf1d3324459f34658cb8957b51 pmc_31784545.pdf Title: First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity Subject: Scientific Reports, doi:10.1038/s41598-019-54042-z Keywords: Author: Alexander W. A. Kellner Creator: Springer Producer: iText® 5.3.5 ©2000-2012 1T3XT BVBA (AGPL-version) CreationDate: Sat Nov 23 10:35:38 2019 CET ModDate: Mon Nov 25 08:55:56 2019 CET Tagged: no UserProperties: no Suspects: no Form: AcroForm JavaScript: no Pages: 9 Encrypted: no Page size: 595.276 x 782.362 pts Page rot: 0 File size: 4563568 bytes Optimized: no PDF version: 1.4 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- QNDYFR+Corbel-Bold CID TrueType Identity-H yes yes yes 26 0 BLQPWL+Corbel-BoldItalic Type 1C WinAnsi yes yes yes 393 0 MinionPro-Regular Type 1C Custom yes no yes 28 0 UKFGGJ+Corbel-Bold2 Type 1C WinAnsi yes yes yes 29 0 MinionPro-It Type 1C Custom yes no yes 31 0 ARGAMQ+Corbel2 Type 1C WinAnsi yes yes yes 32 0 TZQVVL+Corbel CID TrueType Identity-H yes yes yes 33 0 MyriadPro-It Type 1C WinAnsi yes no yes 34 0 MinionPro-Bold Type 1C WinAnsi yes no yes 27 0 EuclidSymbol Type 1C Custom yes no yes 30 0 WVAMGF+TimesNewRomanPSMT Type 1C WinAnsi yes yes yes 273 0 Jhove (Rel. 1.22.1, 2019-04-17) Date: 2020-11-19 06:07:41 CET RepresentationInformation: pmc_31784545.pdf ReportingModule: PDF-hul, Rel. 1.12.1 (2019-04-17) LastModified: 2020-08-03 03:38:02 CEST Size: 4563568 Format: PDF Version: 1.4 Status: Well-Formed, but not valid SignatureMatches: PDF-hul ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 4. Reck, H. Die deutschostafrikanischen Flugsaurier. Zentralblatt fu͜戀爀 䴀椀渀攀爀愀氀漀最椀攀Ⰰ 䜀攀漀氀漀最椀攀 甀渀搀 倀愀氀愃漀渀琀漀氀漀最椀攀⸀⃾＀䄀戀琀⸀ 䋾＀⃾＀㟾＀Ⰰ ㌀㈀ㄠጀ㌀㌀㘀 ㄀㤀㌀㄀⻾＀㨀㌀㈩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㐳ㄮ㈸㈠㈹㈮㌵㜠㐳㐮ㄷ㈠㈸〮㈱㙝⽈⽎⽁值㰯丠㌷㐠〠刾㸾㸊敮摯扪ਲ਼㜴‰扪਼㰯呹灥⽘佢橥捴⽒敳潵牣敳㰼㸾⽓畢瑹灥⽆潲洯䉂潸嬰⸰‰⸰′⸸㠹㤸‱⸰崯䱥湧瑨‱〯䙩汴敲⽆污瑥䑥捯摥㸾獴牥慭ੈ褂࠰Ċ敮摳瑲敡洊敮摯扪ਲ਼㔱‰扪਼㰯䘠㐯呹灥⽁湮潴⽄敳琨SpringerNature_SciRep_54042.indd: 6. Costa, F. R., Sayão, J. M. & Kellner, A. W. A. New pterosaur material from the Upper Jurassic of Tendaguru (Tanzania) Africa. Hist. Biol. 27, 646–655 (2015). :33' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 1. Geographical location where the new pterosaur, Mimodactylus libanensis gen. et sp. nov., was collected. (a) Geological map of Lebanon (adapted from Dubertret14). (b) Detail showing the location of the most important fossil L:39' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 14. Dubertret, L. Carte géologique du Liban au 200 000e. Ministère des Travaux publics, Beyrouth (1955). :44' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 2. Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the hu:52' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 3. Mimodactylus libanensis gen. et sp. nov. (a) Skull and lower jaw. (b) Detail of the dentition. Scale bars, a: 10⁜琀洀洀Ⰰ 戀㨀 ㄠ洀洀⸀ 䄀戀戀爀攀瘀椀愀琀椀漀渀猀⸀ 栀礀㨀 栀礀漀椀搀㬀 洀愀渀㨀 洀愀渀搀椀戀氀攀㬀 漀搀瀀㨀 漀搀漀渀琀漀椀搀 瀀爀漀挀攀猀猀㬀 瀀氀㨀 瀀愀氀愀琀椀渀攀㬀 瀀洀㨀 瀀爀攀洀愀砀椀氀氀愀㬀 瀀瀀氀昀㨀 瀀猀琀㨀㔀㐩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㔰㘮㤶㜠㌷㐮㜶㌠㔱ㄮ㌵㜠㌶㈮㘲㉝⽈⽎⽁值㰯丠㠸‰⁒㸾㸾੥湤潢樊㔸‰扪਼㰯䘠㐯呹灥⽁湮潴⽄敳琨SpringerNature_SciRep_54042.indd: 21. Andres, B. & Qiang, J. Istiodactylus (Pterosauria, Pterodactyloidea) from the Lower Cretaceous of Liaoning, China. J. Vert. Paleontol. 26, 70–78 (2006). :55' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 3. Mimodactylus libanensis gen. et sp. nov. (a) Skull and lower jaw. (b) Detail of the dentition. Scale bars, a: 10⁜琀洀洀Ⰰ 戀㨀 ㄠ洀洀⸀ 䄀戀戀爀攀瘀椀愀琀椀漀渀猀⸀ 栀礀㨀 栀礀漀椀搀㬀 洀愀渀㨀 洀愀渀搀椀戀氀攀㬀 漀搀瀀㨀 漀搀漀渀琀漀椀搀 瀀爀漀挀攀猀猀㬀 瀀氀㨀 瀀愀氀愀琀椀渀攀㬀 瀀洀㨀 瀀爀攀洀愀砀椀氀氀愀㬀 瀀瀀氀昀㨀 瀀猀㨀㘀ㄩ⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈷㌮ㄷ㘠㌴㐮㠸′㠱⸳㠷″㌲⸶㈲崯䠯丯䅐㰼⽎‹㔠〠刾㸾㸊敮摯扪ਸ਼㔠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾＀￾＀㈀㘀⻾￾＀￾￾￾￾＀圀愀渀最Ⰰ 堀⸀ ☀ 䰀ﰀⰀ 䨀⸀ 䐀椀猀挀漀瘀攀爀礀 漀昀 愀 瀀琀攀爀漀搀愀挀琀礀氀椀搀 瀀琀攀爀漀猀愀甀爀 昀爀漀洀 琀栀攀 夀椀砀椀愀渀 䘀漀爀洀愀琀椀漀渀 漀昀 眀攀猀琀攀爀渀 䰀椀愀漀渀椀渀最Ⰰ 䌀栀椀渀愀⸀⃾＀䌀栀椀渀攀猀攀 匀挀椀⸀ 䈀甀氀氀⻾＀⃾＀㐀㛾＀Ⰰ ㄠጀ㘀 ㈀　　㄀⻾＀㨀㘀㈩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㌶〮〷㐠㌴㐮㠸″㘵⸶㔱″㌲⸶㈲崯䠯丯䅐㰼⽎‹㘠〠刾㸾㸊敮摯扪ਸ਼㘠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾＀￾＀㈀㜀⻾￾＀￾￾￾￾＀刀漀搀爀椀最甀攀猀Ⰰ 吀⸀Ⰰ 䨀椀愀渀最Ⰰ 匀⸀Ⰰ 䌀栀攀渀最Ⰰ 堀⸀Ⰰ 圀愀渀最Ⰰ 堀⸀ ☀ 䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀ 䄀 渀攀眀 琀漀漀琀栀攀搀 瀀琀攀爀愀渀漀搀漀渀琀漀椀搀 倀琀攀爀漀猀愀甀爀椀愀Ⰰ 倀琀攀爀漀搀愀挀琀礀氀漀椀搀攀愀 昀爀漀洀 琀栀攀 䨀椀甀昀漀琀愀渀最 䘀漀爀洀愀琀椀漀渀 䰀漀眀攀爀 䌀爀攀琀愀挀攀漀甀猀Ⰰ 䄀瀀琀椀愀渀 漀昀 䌀栀椀渀愀 愀渀搀 挀漀洀洀攀渀琀猀 漀渀⃾＀䰀椀愀漀渀椀渀最漀瀀琀攀爀甀猀 最甀槾＀ 圀愀渀最 愀渀搀 娀㨀㘀㌩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㐳㐮㐱㘠㌴㐮㠸‴㌹⸹㤲″㌲⸶㈲崯䠯丯䅐㰼⽎‹㜠〠刾㸾㸊敮摯扪ਸ਼㜠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾￾￾￾￾￾￾￾￾＀䘀椀最甀爀攀 ㏾￾＀⻾￾＠˾￾￾￾＀䴀椀洀漀搀愀挀琀礀氀甀猀 氀椀戀愀渀攀渀猀椀現＀ 最攀渀⸀ 攀琀 猀瀀⸀ 渀漀瘀⸀ ＀懾＀ 匀欀甀氀氀 愀渀搀 氀漀眀攀爀 樀愀眀⸀ ＀拾＀ 䐀攀琀愀椀氀 漀昀 琀栀攀 搀攀渀琀椀琀椀漀渀⸀ 匀挀愀氀攀 戀愀爀猀Ⰰ 愀㨀 ㄀〠洀洀Ⰰ 戀㨀 ㄠ洀洀⸀ 䄀戀戀爀攀瘀椀愀琀椀漀渀猀⸀ 栀礀㨀 挀攀爀愀琀漀戀爀愀渀挀栀椀愀氀 䤀 漀昀 琀栀攀 栀礀漀椀搀 愀瀀瀀愀爀愀琀甀猀㬀 洀愀渀㨀 洀愀渀搀椀戀氀攀㬀 漀搀瀀㨀 漀搀漀渀琀漀椀搀 瀀爀漀挀攀猀㨀㈀㄀ㄩ⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈶㌮㜲㈠㌳㐮㠸′㜲⸶㐲″㈲⸶㈲崯䠯丯䅐㰼⽎‹㠠〠刾㸾㸊敮摯扪ਸ਼㠠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾＀￾＀㈀㠀⻾￾＀￾￾￾￾＀䰀ﰀⰀ 䨀⸀Ⰰ 堀甀Ⰰ 䰀⸀ ☀ 䨀椀Ⰰ 儀⸀ 刀攀猀琀甀搀礀 漀昀⃾＀䰀椀愀漀砀椀瀀琀攀爀甀現＀ 䤀猀琀椀漀搀愀挀琀礀氀椀搀愀攀㨀 倀琀攀爀漀猀愀甀爀椀愀Ⰰ 眀椀琀栀 挀漀洀洀攀渀琀猀 漀渀 琀栀攀 䌀栀椀渀攀猀攀 椀猀琀椀漀搀愀挀琀礀氀椀搀猀 瀀琀攀爀漀猀愀甀爀猀⸀⃾＀娀椀琀琀攀氀椀愀渀懾＀⃾＀䈀㈀㣾＀Ⰰ ㈀㈀㤠ጀ㈀㐀㄀ ㈀　　㠀⻾＀㨀㘀㔩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈵ㄮ〷㠠㌲㐮㜶㌠㈵㘮㜱ㄠ㌱㈮㘲㉝⽈⽎⽁值㰯丠㤹‰⁒㸾㸾੥湤潢樊㘹‰扪਼㰯䘠㐯呹灥⽁湮潴⽄敳琨SpringerNature_SciRep_54042.indd: 21. Andres, B. & Qiang, J. Istiodactylus (Pterosauria, Pterodactyloidea) from the Lower Cretaceous of Liaoning, China. J. Vert. Paleontol. 26, 70–78 (2006). :66' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 3. Mimodactylus libanensis gen. et sp. nov. (a) Skull and lower jaw. (b) Detail of the dentition. Scale bars, a: 10⁜琀洀洀Ⰰ 戀㨀 ㄠ洀洀⸀ 䄀戀戀爀攀瘀椀愀琀椀漀渀猀⸀ 栀礀㨀 挀攀爀愀琀漀戀爀愀渀挀栀椀愀氀 䤀 漀昀 琀栀攀 栀礀漀椀搀 愀瀀瀀愀爀愀琀甀猀㬀 洀愀渀㨀 洀愀渀搀椀戀氀攀㬀 漀搀瀀㨀 漀搀漀渀琀漀椀搀 瀀爀漀挀攀猀㨀㈀㄀ㄩ⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈶㌮㜲㈠㌳㐮㠸′㜲⸶㐲″㈲⸶㈲崯䠯丯䅐㰼⽎‹㠠〠刾㸾㸊敮摯扪ਸ਼㠠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾＀￾＀㈀㠀⻾￾＀￾￾￾￾＀䰀ﰀⰀ 䨀⸀Ⰰ 堀甀Ⰰ 䰀⸀ ☀ 䨀椀Ⰰ 儀⸀ 刀攀猀琀甀搀礀 漀昀⃾＀䰀椀愀漀砀椀瀀琀攀爀甀現＀ 䤀猀琀椀漀搀愀挀琀礀氀椀搀愀攀㨀 倀琀攀爀漀猀愀甀爀椀愀Ⰰ 眀椀琀栀 挀漀洀洀攀渀琀猀 漀渀 琀栀攀 䌀栀椀渀攀猀攀 椀猀琀椀漀搀愀挀琀礀氀椀搀猀 瀀琀攀爀漀猀愀甀爀猀⸀⃾＀娀椀琀琀攀氀椀愀渀懾＀⃾＀䈀㈀㣾＀Ⰰ ㈀㈀㤠ጀ㈀㐀㄀ ㈀　　㠀⻾＀㨀㘀㔩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈵ㄮ〷㠠㌲㐮㜶㌠㈵㘮㜱ㄠ㌱㈮㘲㉝⽈⽎⽁值㰯丠㤹‰⁒㸾㸾੥湤潢樊㘹‰扪਼㰯䘠㐯呹灥⽁湮潴⽄敳琨SpringerNature_SciRep_54042.indd: 21. Andres, B. & Qiang, J. Istiodactylus (Pterosauria, Pterodactyloidea) from the Lower Cretaceous of Liaoning, China. J. Vert. Paleontol. 26, 70–78 (2006). :66' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 21. Andres, B. & Qiang, J. Istiodactylus (Pterosauria, Pterodactyloidea) from the Lower Cretaceous of Liaoning, China. J. Vert. Paleontol. 26, 70–78 (2006). :66' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 25. Wang, X., Campos, D. A., Zhou, Z. & Kellner, A. W. A. A primitive istiodactylid pterosaur (Pterodactyloidea) from the Jiufotang Formation (Early Cretaceous), northeast China. Zootaxa 1813, 1–18 (2008). :68' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 23. Martill, D. M. A functional odontoid in the dentary of the Early Cretaceous pterosaur Istiodactylus latidens: Implications for feeding. Cretac. Res. 47, 56–65 (2014). :69' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 2. Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the h:72' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 2. Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the :73' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 18. Kellner, A. W. A. Pterosaur phylogeny and comments on the evolutionary history of the group. Geol. Soc. London Spec. Publ. 217(1), 105–137 (2003). :74' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 2. Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the:76' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 2. Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of th:79' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 3. Mimodactylus libanensis gen. et sp. nov. (a) Skull and lower jaw. (b) Detail of the dentition. Scale bars, a: 10⁜琀洀洀Ⰰ 戀㨀 ㄠ洀洀⸀ 䄀戀戀爀攀瘀椀愀琀椀漀渀猀⸀ 栀礀㨀 栀礀漀椀搀㬀 洀愀渀㨀 洀愀渀搀椀戀氀攀㬀 漀搀瀀㨀 漀搀漀渀琀漀椀搀 瀀爀漀挀攀猀猀㬀 瀀氀㨀 瀀愀氀愀琀椀渀攀㬀 瀀洀㨀 瀀爀攀洀愀砀椀氀氀愀㬀 瀀瀀氀昀㨀 㨀㠀㌩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㐰㐮㐹ㄠㄷ㈮㌸‴ㄲ⸷㘲‱㘰⸱㈲崯䠯丯䅐㰼⽎‱㌷‰⁒㸾㸾੥湤潢樊ㄲ㘠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾￾￾￾＀￾＀㈀㔀⻾￾＀￾￾￾￾＀圀愀渀最Ⰰ 堀⸀Ⰰ 䌀愀洀瀀漀猀Ⰰ 䐀⸀ 䄀⸀Ⰰ 娀栀漀甀Ⰰ 娀⸀ ☀ 䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀ 䄀 瀀爀椀洀椀琀椀瘀攀 椀猀琀椀漀搀愀挀琀礀氀椀搀 瀀琀攀爀漀猀愀甀爀 倀琀攀爀漀搀愀挀琀礀氀漀椀搀攀愀 昀爀漀洀 琀栀攀 䨀椀甀昀漀琀愀渀最 䘀漀爀洀愀琀椀漀渀 䔀愀爀氀礀 䌀爀攀琀愀挀攀漀甀猀Ⰰ 渀漀爀琀栀攀愀猀琀 䌀栀椀渀愀⸀⃾＀娀漀漀琀愀砀懾＀⃾＀㄀㠀㄀㏾＀Ⰰ ㄠጀ㄀㠀 ㈀　　㠀⻾＀㨀㠀㐩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈷㜮㠷㐠ㄶ㈮㌸′㠳⸶〳‱㔰⸱㈲崯䠯丯䅐㰼⽎‱㌸‰⁒㸾㸾੥湤潢樊ㄲ㜠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾￾＀￾＀㈀㘀⻾￾＀￾￾￾￾＀圀愀渀最Ⰰ 堀⸀ ☀ 䰀ﰀⰀ 䨀⸀ 䐀椀猀挀漀瘀攀爀礀 漀昀 愀 瀀琀攀爀漀搀愀挀琀礀氀椀搀 瀀琀攀爀漀猀愀甀爀 昀爀漀洀 琀栀攀 夀椀砀椀愀渀 䘀漀爀洀愀琀椀漀渀 漀昀 眀攀猀琀攀爀渀 䰀椀愀漀渀椀渀最Ⰰ 䌀栀椀渀愀⸀⃾＀䌀栀椀渀攀猀攀 匀挀椀⸀ 䈀甀氀氀⻾＀⃾＀㐀㛾＀Ⰰ ㄠጀ㘀 ㈀　　㄀⻾＀㨀㠀㔩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㌵㔮㈵㈠ㄶ㈮㌸″㘰⸹㠱‱㔰⸱㈲崯䠯丯䅐㰼⽎‱㌹‰⁒㸾㸾੥湤潢樊ㄲ㠠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾＀￾＀㈀㤀⻾￾＀￾￾￾￾＀刀漀搀爀椀最甀攀猀Ⰰ 吀⸀ ☀ 䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀ 吀愀砀漀渀漀洀椀挀 爀攀瘀椀攀眀 漀昀 琀栀攀⃾＀伀爀渀椀琀栀漀挀栀攀椀爀甀現＀ 挀漀洀瀀氀攀砀 倀琀攀爀漀猀愀甀爀椀愀 昀爀漀洀 琀栀攀 䌀爀攀琀愀挀攀漀甀猀 漀昀 䔀渀最氀愀渀搀⸀⃾＀娀漀漀䬀攀礀現＀⃾＀㌀　㣾＀Ⰰ ㄠጀ㄀㄀㈀ ㈀　㄀㌀⻾＀㨀㠀㘩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㐰㤮㘹㈠ㄴ㈮㌸‴ㄵ⸳〸‱㌰⸱㈲崯䠯丯䅐㰼⽎‱㐰‰⁒㸾㸾੥湤潢樊ㄲ㤠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾＀￾＀㈀　⻾￾＀￾￾￾￾＀䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀ 䌀漀洀洀攀渀琀猀 漀渀 吀爀椀愀猀猀椀挀 瀀琀攀爀漀猀愀甀爀猀 眀椀琀栀 搀椀猀挀甀猀猀椀漀渀 愀戀漀甀琀 漀渀琀漀最攀渀礀 愀渀搀 搀攀猀挀爀椀瀀琀椀漀渀 漀昀 渀攀眀 琀愀砀愀⸀⃾＀䄀渀⸀ 䄀挀愀搀⸀ 䈀爀愀猀⸀ 䌀椀渀挀⻾＀⃾＀㠀㟾＀Ⰰ 㘀㘀㤠ጀ㘀㠀㤀 ㈀　㄀㔀⻾＀㨀㠀㜩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈶㔮㠳㠠㄰㈮㌸′㜱⸴㤳‹〮ㄲ㈲崯䠯丯䅐㰼⽎‱㐱‰⁒㸾㸾੥湤潢樊ㄳ〠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾＀￾＀㈀㐀⻾￾＀￾￾￾￾＀圀愀渀最Ⰰ 堀⸀Ⰰ 䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀Ⰰ 娀栀漀甀Ⰰ 娀⸀ ☀ 䌀愀洀瀀漀猀Ⰰ 䐀⸀ 䄀⸀ 倀琀攀爀漀猀愀甀爀 搀椀瘀攀爀猀椀琀礀 愀渀搀 昀愀甀渀愀氀 琀甀爀渀漀瘀攀爀 椀渀 䌀爀攀琀愀挀攀漀甀猀 琀攀爀爀攀猀琀爀椀愀氀 攀挀漀猀礀猀琀攀洀猀 椀渀 䌀栀椀渀愀⸀⃾＀一愀琀甀爀旾＀⃾＀㐀㌀㟾＀Ⰰ 㠀㜀㔠ጀ㠀㜀㤀 ㈀　　㔀⻾＀㨀㠀㠩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㈷㐮㔵㐠㄰㈮㌸′㠰⸲〸‹〮ㄲ㈲崯䠯丯䅐㰼⽎‱㐲‰⁒㸾㸾੥湤潢樊ㄳㄠ〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾￾￾＀￾＀㈀㄀⻾￾＀￾￾￾￾＀䄀渀搀爀攀猀Ⰰ 䈀⸀ ☀ 儀椀愀渀最Ⰰ 䨀⸀⃾＀䤀猀琀椀漀搀愀挀琀礀氀甀現＀ 倀琀攀爀漀猀愀甀爀椀愀Ⰰ 倀琀攀爀漀搀愀挀琀礀氀漀椀搀攀愀 昀爀漀洀 琀栀攀 䰀漀眀攀爀 䌀爀攀琀愀挀攀漀甀猀 漀昀 䰀椀愀漀渀椀渀最Ⰰ 䌀栀椀渀愀⸀⃾＀䨀⸀ 嘀攀爀琀⸀ 倀愀氀攀漀渀琀漀氀⻾＀⃾＀㈀㛾＀Ⰰ 㜀〠ጀ㜀㠀 ㈀　　㘀⻾＀㨀㠀㤩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㔳ㄮ㐷㐠㄰㈮㌸‵㌷⸱㈹‹〮ㄲ㈲崯䠯丯䅐㰼⽎‱㐳‰⁒㸾㸾੥湤潢樊ㄳ㈠〠潢樊㰼⽆‴⽔祰支䅮湯琯䑥獴⣾＀匀瀀爀椀渀最攀爀一愀琀甀爀攀开匀挀椀刀攀瀀开㔀㐀　㐀㈀⸀椀渀搀搀㫾￾￾￾＀￾＀㄀㠀⻾￾＀￾￾￾￾＀䬀攀氀氀渀攀爀Ⰰ 䄀⸀ 圀⸀ 䄀⸀ 倀琀攀爀漀猀愀甀爀 瀀栀礀氀漀最攀渀礀 愀渀搀 挀漀洀洀攀渀琀猀 漀渀 琀栀攀 攀瘀漀氀甀琀椀漀渀愀爀礀 栀椀猀琀漀爀礀 漀昀 琀栀攀 最爀漀甀瀀⸀⃾＀䜀攀漀氀⸀ 匀漀挀⸀ 䰀漀渀搀漀渀 匀瀀攀挀⸀ 倀甀戀氀⻾＀⃾＀㈀㄀㟾＀㄀Ⰰ ㄀　㔠ጀ㄀㌀㜀 ㈀　　㌀⻾＀㨀㤀〩⽂潲摥牛〠〠そ⽓畢瑹灥⽌楮欯䉓㰼⽗‰⽔祰支䉯牤敲⽓⽓㸾⽒散瑛㔴㤮〲㈠㘲⸲㘳㈠㔵㐮㘳㠠㔰⸱㈲㉝⽈⽎⽁值㰯丠ㄴ㐠〠刾㸾㸊敮摯扪਱㌳‰扪਼㰯䘠㐯呹灥⽁湮潴⽄敳琨SpringerNature_SciRep_54042.indd: 34. Holgado, B. et al. On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria. Sci. Rep. 9, 4940 (2019). :99' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 26. Wang, X. & Lü, J. Discovery of a pterodactylid pterosaur from the Yixian Formation of western Liaoning, China. Chinese Sci. Bull. 46, 1–6 (2001). :85' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 24. Wang, X., Kellner, A. W. A., Zhou, Z. & Campos, D. A. Pterosaur diversity and faunal turnover in Cretaceous terrestrial ecosystems in China. Nature 437, 875–879 (2005). :91' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 4. An overview of Mimodactylus libanensis gen. et sp. nov. (a) Phylogenetic relationships of Mimodactylus libanensis gen. et sp. nov. within Ornithocheiroidea. Colours show their continental origin: Africa (brown), Asia (orange), Eu:98' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 30. Wellnhofer, P. The illustrated encyclopedia of pterosaurs. London: Salamander Books. 192 pp. (1991). :101' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 40. Wang, X., Kellner, A. W. A., Jiang, S. & Cheng, X. New toothed flying reptile from Asia: close similarities between early Cretaceous pterosaur faunas from China and Brazil. Sci. Nat. 99, 249–257 (2012). :107' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 41. Wang, X., Rodrigues, T., Jiang, S., Cheng, X. & Kellner, A. W. A. An Early Cretaceous pterosaur with an unusual mandibular crest from China and a potential novel feeding strategy. Sci. Rep. 4, 6329 (2014). :108' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 42. Frey, E., Buchy, M. C., Stinnesbeck, W., González, A. G. & Stefano, A. Muzquizopteryx coahuilensis n.g., n.sp., a nyctosaurid pterosaur with soft tissue preservation from the Coniacian (Late Cretaceous) of northeast Mexico (Coahuila). Or:110' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 43. Wang, X. & Zhou, Z. Pterosaur assemblages of the Jehol Biota and their implication for the Early Cretaceous pterosaur radiation. Geol. J. 41, 405–418 (2006). :111' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 44. Kellner, A. W. A. & Campos, D. A. The function of the cranial crest and jaws of a unique pterosaur from the Early Cretaceous of Brazil. Science 297, 389–392 (2002). :112' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 45. Wellnhofer, P. & Kellner, A. W. A. The skull of Tapejara wellnhoferi Kellner (Reptilia, Pterosauria) from the Lower Cretaceous Santana Formation of the Araripe Basin, Northeastern Brazil. Mitt. Bayer. Staatssamml. Paläontol. Hist. Geol.:113' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 46. Vullo, R. et al. A New Crested Pterosaur from the Early Cretaceous of Spain: The First European Tapejarid (Pterodactyloidea: Azhdarchoidea). PLoS One 7, e38900 (2012). :114' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 47. Wang, X., Kellner, A. W. A., Zhou, Z. & Campos, D. A. Discovery of a rare arboreal forest-dwelling flying reptile (Pterosauria, Pterodactyloidea) from China. Proc. Natl. Acad. Sci. USA 105, 1983–1987 (2008). :116' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 48. Kellner, A. W. A. & Calvo, J. O. New azhdarchoid pterosaur (Pterosauria, Pterodactyloidea) with an unusual lower jaw from the Portezuelo Formation (Upper Cretaceous), Neuquén Group, Patagonia, Argentina. An. Acad. Bras. Ciênc. 89, 2003–:119' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 49. Witton, M. P. & Naish, D. A. Reappraisal of azhdarchid pterosaur functional morphology and paleoecology. PLoS One 3, e2271 (2008). :120' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 50. Edmund, A. G. Dentition. In: Biology of the Reptilia (eds Gans, C., Bellairs, A. A. & Parsons, N.), 117–200 (Academic Press, London, 1969). :121' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 53. Evans, A. R. & Sanson, G. D. The effect of tooth shape on the breakdown of insects. J. Zool. 246, 391–400 (1998). :122' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 54. Bennett, S. C. A second specimen of the pterosaur Anurognathus ammoni. Paläont. Z. 81, 376–398 (2007). :124' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 55. Wilson, D. S. The adequacy of body size as a niche difference. Am. Nat. 109(970), 769–784 (1975). :125' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 57. Dial, K. P. Evolution of avian locomotion: correlates of flight style, locomotor modules, nesting biology, body size, development, and the origin of flapping flight. Auk 120(4), 941–952 (2003). :126' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 56. Barclay, R. M. R. & Brigham, R. M. Prey detection, dietary niche breadth, and body size in bats: why are aerial insectivorous bats so small? Am. Nat. 137(5), 693–703 (1991). :127' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 58. Witton, M. P. & Habib, M. B. On the size and flight diversity of giant pterosaurs, the use of birds as pterosaur analogues and comments on pterosaur flightlessness. PLoS One 5, e13982 (2010). :128' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 4. An overview of Mimodactylus libanensis gen. et sp. nov. (a) Phylogenetic relationships of Mimodactylus libanensis gen. et sp. nov. within Ornithocheiroidea. Colours show their continental origin: Africa (brown), Asia (orange), E:129' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 59. Xavier, J. C., Croxall, J. P., Trathan, P. N. & Wood, A. G. Feeding strategies and diets of breeding grey-headed and wandering albatrosses at South Georgia. Marine Biology 143, 221–232 (2003). :132' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd:Figure 5. Life reconstruction of Mimodactylus libanensis gen. et sp. nov. Artwork of Julius T. Csotonyi.:142' cannot be found Offset: 4040093 ID: PDF-HUL-122 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'SpringerNature_SciRep_54042.indd: 34. Holgado, B. et al. On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria. Sci. Rep. 9, 4940 (2019). :144' cannot be found Offset: 4040093 MIMEtype: application/pdf PDFMetadata: Objects: 480 FreeObjects: 1 IncrementalUpdates: 0 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseOutlines Language: EN Outlines: Item: Title: First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity Children: Item: Title: Results Children: Item: Title: Systematic palaeontology. Item: Title: Branch-based definition. Item: Title: Diagnosis. Item: Title: Included taxa. Item: Title: Branch-based definition. Item: Title: Diagnosis. Item: Title: Included species. Item: Title: Etymology. Item: Title: Holotype. Item: Title: Locality and horizon. Item: Title: Diagnosis. Item: Title: Comparative description. Item: Title: Discussion Item: Title: Methods Children: Item: Title: Nomenclatural acts. Item: Title: Phylogenetic analysis. Item: Title: Acknowledgements Item: Title: Figure 1 Geographical location where the new pterosaur, Mimodactylus libanensis gen. Item: Title: Figure 2 Mimodactylus libanensis gen. Item: Title: Figure 3 Mimodactylus libanensis gen. Item: Title: Figure 4 An overview of Mimodactylus libanensis gen. Item: Title: Figure 5 Life reconstruction of Mimodactylus libanensis gen. Info: Title: First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity Author: Alexander W. A. Kellner Subject: Scientific Reports, doi:10.1038/s41598-019-54042-z Keywords: Creator: Springer Producer: iText® 5.3.5 ©2000-2012 1T3XT BVBA (AGPL-version) CreationDate: Sat Nov 23 11:05:38 CET 2019 ModDate: Mon Nov 25 08:55:56 CET 2019 ID: 0x4010aa9435c37f1ece61bd7b31bff4fa, 0xdaf585d53a56030cb794d480455995f1 Filters: FilterPipeline: FlateDecode FilterPipeline: DCTDecode Images: Image: NisoImageMetadata: FormatName: image/png ImageWidth: 1900 ImageHeight: 1531 BitsPerSample: 8 BitsPerSampleUnit: integer Filter: FlateDecode Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 1650 ImageHeight: 1443 BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 1350 ImageHeight: 743 BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/png ImageWidth: 1900 ImageHeight: 1586 BitsPerSample: 8 BitsPerSampleUnit: integer Filter: FlateDecode Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 1500 ImageHeight: 1500 BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/png ImageWidth: 1100 ImageHeight: 90 ColorSpace: black is zero BitsPerSample: 1 BitsPerSampleUnit: integer Filter: FlateDecode Intent: RelativeColorimetric Name: X Fonts: Type0: Font: BaseFont: TZQVVL+Corbel Encoding: Identity-H ToUnicode: true Font: BaseFont: QNDYFR+Corbel-Bold Encoding: Identity-H ToUnicode: true Type1: Font: BaseFont: ARGAMQ+Corbel2 FontSubset: true FirstChar: 0 LastChar: 255 FontDescriptor: FontName: ARGAMQ+Corbel2 Flags: Nonsymbolic FontBBox: -553, -239, 1110, 952 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: WVAMGF+TimesNewRomanPSMT FontSubset: true FirstChar: 32 LastChar: 120 FontDescriptor: FontName: WVAMGF+TimesNewRomanPSMT Flags: Serif, Nonsymbolic FontBBox: -568, -307, 2000, 1007 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: MyriadPro-It FirstChar: 0 LastChar: 255 FontDescriptor: FontName: MyriadPro-It Flags: Nonsymbolic, Italic FontBBox: -185, -250, 1090, 953 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: BLQPWL+Corbel-BoldItalic FontSubset: true FirstChar: 0 LastChar: 255 FontDescriptor: FontName: BLQPWL+Corbel-BoldItalic Flags: Nonsymbolic, Italic FontBBox: -494, -239, 1194, 952 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: MinionPro-Bold FirstChar: 0 LastChar: 255 FontDescriptor: FontName: MinionPro-Bold Flags: Serif, Nonsymbolic FontBBox: -319, -360, 1684, 1024 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: MinionPro-Regular FirstChar: 0 LastChar: 255 FontDescriptor: FontName: MinionPro-Regular Flags: Serif, Nonsymbolic FontBBox: -290, -360, 1684, 989 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: UKFGGJ+Corbel-Bold2 FontSubset: true FirstChar: 0 LastChar: 255 FontDescriptor: FontName: UKFGGJ+Corbel-Bold2 Flags: Nonsymbolic FontBBox: -556, -239, 1155, 952 FontFile3: true Encoding: WinAnsiEncoding ToUnicode: true Font: BaseFont: EuclidSymbol FirstChar: 0 LastChar: 255 FontDescriptor: FontName: EuclidSymbol Flags: Symbolic FontBBox: -166, -253, 1090, 950 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: MinionPro-It FirstChar: 0 LastChar: 255 FontDescriptor: FontName: MinionPro-It Flags: Serif, Nonsymbolic, Italic FontBBox: -201, -360, 1684, 1002 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true CIDFontType2: Font: BaseFont: QNDYFR+Corbel-Bold CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: QNDYFR+Corbel-Bold Flags: Symbolic FontBBox: -556, -239, 1155, 952 FontFile2: true Font: BaseFont: TZQVVL+Corbel CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: TZQVVL+Corbel Flags: Symbolic FontBBox: -553, -239, 1110, 952 FontFile2: true XMP: application/pdf doi:10.1038/s41598-019-54042-z Springer US Scientific Reports, doi:10.1038/s41598-019-54042-z First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity Alexander W. A. Kellner Michael W. Caldwell Borja Holgado Fabio M. Dalla Vecchia Roy Nohra Juliana M. Sayamp#x000E3;o Philip J. Currie 10.1038/s41598-019-54042-z http://dx.doi.org/10.1038/s41598-019-54042-z journal Scientific Reports © 2019, The Author(s) 2045-2322 10.1038/s41598-019-54042-z Springer 2019-11-25T08:55:56+01:00 2019-11-23T15:05:38+05:30 2019-11-25T08:55:56+01:00 True iText® 5.3.5 ©2000-2012 1T3XT BVBA (AGPL-version) VoR uuid:ee2b5fec-5e62-4cd6-83fe-384d57a48713 uuid:5e60f574-0175-439f-836b-3bbbb8967ce9 default 1 converted uuid:2ce39784-5570-4983-afd4-0a1c4a1900b5 converted to PDF/A-2b pdfToolbox 2019-11-23T15:06:38+05:30 2 B http://ns.adobe.com/xap/1.0/mm/ xmpMM XMP Media Management Schema internal UUID based identifier for specific incarnation of a document InstanceID URI internal The common identifier for all versions and renditions of a document. OriginalDocumentID URI internal A reference to the original document from which this one is derived. 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