e6cbd99f164983f0cb19f2fb453b988b comptes-rendus-palevol2021v20a29.pdf a0d177603f738f377622dcadd19aef1abfbeece6 comptes-rendus-palevol2021v20a29.pdf c398b2e5211d9a86cdaa50aed63dba82032f7ac7367e239f1ce915a421f0adff comptes-rendus-palevol2021v20a29.pdf Title: Ancestral foxes at the gates of Europe: the Pliocene fox from Çalta Subject: English abstract (résumé anglais) The Pliocene record of genus Vulpes Frisch, 1775 in Eurasia is scarce, coming from few sparse localities. The lack of a comprehensive and integrated revision led to the description of numerous different taxa, often only tentatively related to extant species but not with one another. Çalta-1 is an important Pliocene site located in the Anatolian region of Turkey, dated to 4.0 Ma. In the present review, we reappraise the interesting record of Vulpes galatica Ginsburg, 1998. Morphological and morphometric evidence suggests a strong similarity between this taxon and the early Late Pliocene V. beihaiensis Qiu & Tedford, 1990, recovered from the Chinese Yushe Basin. Such evidence favors the parsimonious interpretation of synonymy between the two species, under the name V. beihaiensis. This hypothesis opens a new interpretation on the biogeography of the Pliocene-Early Pleistocene foxes of Eurasia. Vulpes beihaiensis links the Asian and European records, with its affinity to V. ­alopecoides (Del Campana, 1913) and, eventually, to the extant red fox (Vulpes vulpes (Linnaeus, 1758)). French abstract (résumé français) Le signalement pliocène du genre Vulpes Frisch, 1775 en Eurasie est peu abondant, du fait de quelques localités éparses. L’absence d’une révision intégrée et exhaustive a conduit à la description de nombre de taxons souvent reliés, à titre d’essai, à des espèces actuelles, mais non l’un à l’autre. Calta-1 est un important site pliocène localisé dans la région anatolienne de Turquie, daté de 4 Ma. Dans cet article, nous réexaminons l’intéressant signalement de Vulpes galatica Ginsburg, 1998. Des preuves morphologiques et morphométriques suggèrent une importante similitude entre ce taxon et V. ­beihaiensis Qiu & Tedford, 1990 du début du Pliocène supérieur, récolté dans le bassin d’Yushe, en Chine. De telles preuves plaident en faveur de l’interprétation parcimonieuse d’une synonymie entre les deux espèces, sous le nom de V. beihaiensis. Cette hypothèse permet une nouvelle interprétation des renards d’Eurasie au Plio-Pléistocène-Pléistocène inférieur. Vulpes beihaiensis relie les registres européens et asiatiques, avec son affinité à V. alopecoides (Del Campana, 1913) et, éventuellement, au renard rouge actuel (Vulpes vulpes (Linnaeus, 1758)). Author: Saverio BARTOLINI-LUCENTI Creator: Adobe InDesign 16.2 (Windows) Producer: Adobe PDF Library 15.0 CreationDate: Mon Jul 12 15:57:25 2021 CEST ModDate: Mon Jul 12 15:57:28 2021 CEST Custom Metadata: no Metadata Stream: yes Tagged: yes UserProperties: no Suspects: no Form: none JavaScript: no Pages: 10 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 3263426 bytes Optimized: yes PDF version: 1.5 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- OIQBNR+Wingdings-Regular CID TrueType Identity-H yes yes yes 1291 0 YXQJXX+ITCStoneSans-Semibold Type 1C Custom yes yes yes 1294 0 PGPQWP+HelveticaNeue CID TrueType Identity-H yes yes yes 85 0 RBQHYL+ITCStoneSans-SemiboldItalic Type 1C WinAnsi yes yes yes 82 0 YXQJXX+ITCStoneSans-Semibold Type 1C WinAnsi yes yes yes 83 0 BLMKIR+HelveticaNeue TrueType WinAnsi yes yes yes 132 0 PTNQWP+HelveticaNeue-Italic TrueType WinAnsi yes yes yes 116 0 HCMPOF+HelveticaNeue-SC750 TrueType WinAnsi yes yes yes 120 0 CXZLWP+HelveticaNeue-Italic-SC750 TrueType WinAnsi yes yes yes 129 0 XNMHEZ+HelveticaNeue-BoldItalic TrueType WinAnsi yes yes yes 134 0 VKNVCD+HelveticaNeue-Bold TrueType WinAnsi yes yes yes 138 0 LVNNSX+AGaramondPro-Regular Type 1C Custom yes yes yes 149 0 GWSZAH+AGaramondPro-Semibold Type 1C WinAnsi yes yes yes 161 0 EJPIYL+AGaramondPro-Italic Type 1C Custom yes yes yes 162 0 LVNNSX+AGaramondPro-Regular Type 1C WinAnsi yes yes yes 210 0 BLMKIR+HelveticaNeue TrueType WinAnsi yes yes yes 209 0 FLYEMJ+AGaramond-Regular-SC750 Type 1C WinAnsi yes yes yes 281 0 ECKTYL+HelveticaNeue Type 1C WinAnsi yes yes no 265 0 ECKTYL+HelveticaNeue-Italic Type 1C WinAnsi yes yes no 284 0 LVNNSX+AGaramondPro-Regular Type 1C WinAnsi yes yes yes 245 0 BLMKIR+HelveticaNeue TrueType WinAnsi yes yes yes 250 0 DDVNKN+AGaramondPro-SemiboldItalic Type 1C WinAnsi yes yes yes 306 0 NATVUT+AGaramond-Italic-SC750 Type 1C WinAnsi yes yes yes 321 0 ECKTYL+HelveticaNeue Type 1C WinAnsi yes yes no 320 0 ECKTYL+HelveticaNeue Type 1C WinAnsi yes yes no 371 0 ECKTYL+HelveticaNeue Type 1C WinAnsi yes yes no 391 0 ECKTYL+HelveticaNeue-Italic Type 1C WinAnsi yes yes no 403 0 XCWDEZ+AGaramond-Regular Type 1C WinAnsi yes yes yes 418 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2024-10-02 03:48:30 CEST RepresentationInformation: comptes-rendus-palevol2021v20a29.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2023-03-31 11:51:21 CEST Size: 3263426 Format: PDF Version: 1.5 Status: Well-Formed and valid SignatureMatches: PDF-hul MIMEtype: application/pdf Profile: Linearized PDF PDFMetadata: Objects: 1300 FreeObjects: 809 IncrementalUpdates: 1 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Language: fr-FR Info: Author: Saverio BARTOLINI-LUCENTI Subject: English abstract (résumé anglais) The Pliocene record of genus Vulpes Frisch, 1775 in Eurasia is scarce, coming from few spar猀攀 氀漀挀愀氀椀琀椀攀猀⸀ 吀栀攀 氀愀挀欀 漀昀 愀 挀漀洀瀀爀攀栀攀渀猀椀瘀攀 愀渀搀 椀渀琀攀最爀愀琀攀搀 爀攀瘀椀猀椀漀渀 氀攀搀 琀漀 琀栀攀 搀攀猀挀爀椀瀀琀椀漀渀 漀昀 渀甀洀攀爀漀甀猀 搀椀昀昀攀爀攀渀琀 琀愀砀愀Ⰰ 漀昀琀攀渀 漀渀氀 tentatively related to extant species but not with one another. Çalta-1 is an important Pliocene site located in the Anatolian 爀攀最椀漀渀 漀昀 吀甀爀欀攀礀Ⰰ 搀愀琀攀搀 琀漀 㐀⸀　 䴀愀⸀ 䤀渀 琀栀攀 瀀爀攀猀攀渀琀 爀攀瘀椀攀眀Ⰰ 眀攀 爀攀愀瀀瀀爀愀椀猀攀 琀栀攀 椀渀琀攀爀攀猀琀椀渀最 爀攀挀漀爀搀 漀昀 嘀甀氀瀀攀猀 最愀氀愀琀椀挀愀 䜀椀渀猀戀甀爀最Ⰰ ㄀㤀8. Morphological and morphometric evidence suggests a strong similarity between this taxon and the early Late Pliocene V. beiha椀攀渀猀椀猀 儀椀甀ꀀ☀ 吀攀搀昀漀爀搀Ⰰ ㄀㤀㤀　Ⰰ 爀攀挀漀瘀攀爀攀搀 昀爀漀洀 琀栀攀 䌀栀椀渀攀猀攀 夀甀猀栀攀 䈀愀猀椀渀⸀ 匀甀挀栀 攀瘀椀搀攀渀挀攀 昀愀瘀漀爀猀 琀栀攀 瀀愀爀猀椀洀漀渀椀漀甀猀 椀渀琀攀爀瀀爀攀琀愀琀椀漀渀 漀昀 猀礀渀nymy between the two species, under the name V. beihaiensis. This hypothesis opens a new interpretation on the biogeography of 琀栀攀 倀氀椀漀挀攀渀攀ⴀ䔀愀爀氀礀 倀氀攀椀猀琀漀挀攀渀攀 昀漀砀攀猀 漀昀 䔀甀爀愀猀椀愀⸀ 嘀甀氀瀀攀猀 戀攀椀栀愀椀攀渀猀椀猀 氀椀渀欀猀 琀栀攀 䄀猀椀愀渀 愀渀搀 䔀甀爀漀瀀攀愀渀 爀攀挀漀爀搀猀Ⰰ 眀椀琀栀 椀琀猀 愀昀昀椀渀椀琀礀 琀漀 . ­alopecoides (Del Campana, 1913) and, eventually, to the extant red fox (Vulpes vulpes (Linnaeus, 1758)). French abstra挀琀 爀猀甀洀 昀爀愀渀愀椀猀䰀攀 猀椀最渀愀氀攀洀攀渀琀 瀀氀椀漀挀渀攀 搀甀 最攀渀爀攀 嘀甀氀瀀攀猀 䘀爀椀猀挀栀Ⰰ ㄀㜀㜀㔀 攀渀 䔀甀爀愀猀椀攀 攀猀琀 瀀攀甀 愀戀漀渀搀愀渀琀Ⰰ 搀甀 昀愀椀琀 搀攀 焀甀攀氀焀甀攀猀 氀漀挀愀氀椀琀猀 瀀愀爀猀攀猀⸀ 䰠ᤀ愀戀猀攀渀挀攀 搠ᤀ甀渀攀 爀瘀椀猀椀漀渀 椀渀琀最爀攀 攀琀 攀砀栀愀甀猀琀椀瘀攀 愀 挀漀渀搀甀椀琀  氀愀 搀攀猀挀爀椀瀀琀椀漀渀 搀攀 渀漀洀戀爀攀 搀攀 琀愀砀漀渀猀 猀漀甀瘀攀渀琀 爀攀氀椀猀Ⰰ  titre d’essai, à des espèces actuelles, mais non l’un à l’autre. Calta-1 est un important site pliocène localisé dans la régio渀 愀渀愀琀漀氀椀攀渀渀攀 搀攀 吀甀爀焀甀椀攀Ⰰ 搀愀琀 搀攀 㐀ꀀ䴀愀⸀ 䐀愀渀猀 挀攀琀 愀爀琀椀挀氀攀Ⰰ 渀漀甀猀 爀攀砀愀洀椀渀漀渀猀 氠ᤀ椀渀琀爀攀猀猀愀渀琀 猀椀最渀愀氀攀洀攀渀琀 搀攀 嘀甀氀瀀攀猀 最愀氀愀琀椀挀愀 䜀椀渀猀戀甀爀, 1998. Des preuves morphologiques et morphométriques suggèrent une importante similitude entre ce taxon et V. ­beihaiensis Qiuꀀ☀ 吀攀搀昀漀爀搀Ⰰ ㄀㤀㤀　 搀甀 搀戀甀琀 搀甀 倀氀椀漀挀渀攀 猀甀瀀爀椀攀甀爀Ⰰ 爀挀漀氀琀 搀愀渀猀 氀攀 戀愀猀猀椀渀 搠ᤀ夀甀猀栀攀Ⰰ 攀渀 䌀栀椀渀攀⸀ 䐀攀ꀀ琀攀氀氀攀猀 瀀爀攀甀瘀攀猀 瀀氀愀椀搀攀渀琀 攀渀 昀愀瘀攀甀爀de l’interprétation parcimonieuse d’une synonymie entre les deux espèces, sous le nom de V. beihaiensis. Cette hypothèse permet 甀渀攀 渀漀甀瘀攀氀氀攀 椀渀琀攀爀瀀爀琀愀琀椀漀渀 搀攀猀 爀攀渀愀爀搀猀 搠ᤀ䔀甀爀愀猀椀攀 愀甀 倀氀椀漀ⴀ倀氀椀猀琀漀挀渀攀ⴀ倀氀椀猀琀漀挀渀攀 椀渀昀爀椀攀甀爀⸀ 嘀甀氀瀀攀猀 戀攀椀栀愀椀攀渀猀椀猀 爀攀氀椀攀 氀攀猀 爀攀最椀tres européens et asiatiques, avec son affinité à V. alopecoides (Del Campana, 1913) et, éventuellement, au renard rouge actue氀 嘀甀氀瀀攀猀 瘀甀氀瀀攀猀 䰀椀渀渀愀攀甀猀Ⰰ ㄀㜀㔀㠀⸩⽔楴汥⡁湣敳瑲慬⁦潸敳⁡琠瑨攠条瑥猠潦⁅畲潰攺⁴桥⁐汩潣敮攠景砠晲潭⃇慬瑡 Creator: Adobe InDesign 16.2 (Windows) Producer: Adobe PDF Library 15.0 CreationDate: Mon Jul 12 15:57:25 CEST 2021 ModDate: Mon Jul 12 15:57:28 CEST 2021 ID: 0x745dad546b89d1408dae262fe5684b7d, 0x50dfee08f396ff4ba1f9e103febeee43 Filters: FilterPipeline: FlateDecode FilterPipeline: DCTDecode Images: Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 1037 ImageHeight: 813 ColorSpace: CMYK BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 1036 ImageHeight: 813 ColorSpace: CMYK BitsPerSample: 8 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uuid:32c390bd-725f-4089-8fc1-70cf458a6059 adobe:docid:indd:c1ef6c8e-16fc-11de-88e8-a60412db0680 xmp.id:b9bacc12-7abd-0c49-b391-e4dc5dce57ed proof:pdf xmp.iid:35b9faef-c75d-9648-ae54-2baf92054f02 xmp.did:352454aa-6abb-574b-b40d-b427c3859d02 adobe:docid:indd:c1ef6c8e-16fc-11de-88e8-a60412db0680 default converted from application/x-indesign to application/pdf Adobe InDesign 16.2 (Windows) / 2021-07-12T15:57:25+02:00 application/pdf Saverio BARTOLINI-LUCENTI Joan MADURELL-MALAPEIRA © Publications scientifiques du Muséum national d’Histoire naturelle, Académie des sciences, Paris, 2021 — Comptes Rendus Palevol — http://sciencepress.mnhn.fr English abstract (résumé anglais) The Pliocene record of genus Vulpes Frisch, 1775 in Eurasia is scarce, coming from few sparse localities. The lack of a comprehensive and integrated revision led to the description of numerous different taxa, often only tentatively related to extant species but not with one another. Çalta-1 is an important Pliocene site located in the Anatolian region of Turkey, dated to 4.0 Ma. In the present review, we reappraise the interesting record of Vulpes galatica Ginsburg, 1998. Morphological and morphometric evidence suggests a strong similarity between this taxon and the early Late Pliocene V. beihaiensis Qiu & Tedford, 1990, recovered from the Chinese Yushe Basin. Such evidence favors the parsimonious interpretation of synonymy between the two species, under the name V. beihaiensis. This hypothesis opens a new interpretation on the biogeography of the Pliocene-Early Pleistocene foxes of Eurasia. Vulpes beihaiensis links the Asian and European records, with its affinity to V. ­alopecoides (Del Campana, 1913) and, eventually, to the extant red fox (Vulpes vulpes (Linnaeus, 1758)). French abstract (résumé français) Le signalement pliocène du genre Vulpes Frisch, 1775 en Eurasie est peu abondant, du fait de quelques localités éparses. L’absence d’une révision intégrée et exhaustive a conduit à la description de nombre de taxons souvent reliés, à titre d’essai, à des espèces actuelles, mais non l’un à l’autre. Calta-1 est un important site pliocène localisé dans la région anatolienne de Turquie, daté de 4 Ma. Dans cet article, nous réexaminons l’intéressant signalement de Vulpes galatica Ginsburg, 1998. Des preuves morphologiques et morphométriques suggèrent une importante similitude entre ce taxon et V. ­beihaiensis Qiu & Tedford, 1990 du début du Pliocène supérieur, récolté dans le bassin d’Yushe, en Chine. De telles preuves plaident en faveur de l’interprétation parcimonieuse d’une synonymie entre les deux espèces, sous le nom de V. beihaiensis. Cette hypothèse permet une nouvelle interprétation des renards d’Eurasie au Plio-Pléistocène-Pléistocène inférieur. Vulpes beihaiensis relie les registres européens et asiatiques, avec son affinité à V. alopecoides (Del Campana, 1913) et, éventuellement, au renard rouge actuel (Vulpes vulpes (Linnaeus, 1758)). Ancestral foxes at the gates of Europe: the Pliocene fox from Çalta Carnivora Vulpes Pliocene Eurasia biogeography Carnivora Vulpes Pliocène Eurasie biogéographie http://cr-palevol.fr/20/29 True Adobe PDF Library 15.0 False Muséum national d'Histoire naturelle, Publications scientifiques, Palevol Pages: Page: Label: I Page: Label: II Annotations: Annotation: Subtype: Link Rect: 49, 172, 82, 164 Annotation: Subtype: Link Rect: 86, 172, 126, 164 Annotation: Subtype: Link Rect: 130, 172, 166, 164 Annotation: Subtype: Link Rect: 170, 172, 228, 164 Annotation: Subtype: Link Rect: 232, 172, 326, 164 Annotation: Subtype: Link Rect: 330, 172, 356, 164 Annotation: Subtype: Link Rect: 445, 172, 474, 164 Annotation: Subtype: Link Rect: 478, 172, 507, 164 Annotation: Subtype: Link Rect: 510, 172, 542, 164 Annotation: Subtype: Link Rect: 49, 153, 149, 144 Annotation: Subtype: Link Rect: 153, 153, 234, 144 Annotation: Subtype: Link Rect: 238, 153, 326, 144 Annotation: Subtype: Link Rect: 330, 153, 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