Journal of South American Earth Sciences 109 (2021) 103237 Contents lists available at ScienceDirect Journal of South American Earth Sciences journal homepage: www.elsevier.com/locate/jsames Mexican Upper Cretaceous rudists (Hippuritida, Bivalvia): Taxonomic, stratigraphic, and geologic data Jose Maria Pons a, 1, *, Pedro García Barrera b, Angélica Oviedo c, Enric Vicens a a Departament de Geologia, Universitat Autònoma de Barcelona, 08193, Bellaterra, Barcelona, Spain b Museo de Paleontología, Facultad de Ciencias, Universidad Nacional Autónoma de México, Mexico c Facultad de Ingeniería, Universidad Autónoma de Chihuahua, Mexico A R T I C L E I N F O A B S T R A C T Keywords: Analysis of seventy published papers dealing with Mexican Upper Cretaceous rudists, together with consultation Upper cretaceous of the newest geological maps covering the areas with mentioned rudist fossil localities, evidenced that: a) Mexico several among the Turonian-Maastrichtian species mentioned are junior synonymies, nomen nudum, insuffi­ Rudists ciently described, or have untraceable types. According to modern standards, around sixty species seem Geology Stratigraphy acceptable, Antillocaprinidae (12), Hippuritidae (11), Plagioptychidae (9), and Radiolitidae (27). Only three Taxonomy species (Turonian) are common with Eurasia. All the Santonian-Maastrichtian rudists (54) are exclusively American species, either ascribed to Eurasian (9) or to American (21) genera. b) Research on rudists helped to improve the stratigraphy in some areas, and to point out or clear up some geological issues in others. c) The restudy of the geology in some areas, of some rudist collections in museums, and of some rudist groups, are much necessary and must be undertaken. 1. Introduction during the twenty-first century, centered on Mexican material or including it within a wider scope, and paying special attention to The Lower and middle Cretaceous limestones from Mexico have been biostratigraphy. deeply investigated since the end of nineteenth century, due mainly to The scope of this paper is (1) to provide a complete updated survey their economic interest in petroleum exploration; they have also been on the present day knowledge of Mexican Late Cretaceous rudists, (2) to intensely quarried as material for construction. Since rudist bivalve stress their contribution to the stratigraphy and geology of the Mexican shells are a conspicuous component of these limestones, reports or Upper Cretaceous, and (3) to point out those aspects deserving further paleontological studies of this fossil group are quite abundant in the study. geological literature of the area. Although not as conspicuous as Lower This paper is dedicated to the memory of the late Dr. Gloria and middle Cretaceous ones, Upper Cretaceous limestones and their Alencáster. For her important contribution to the knowledge of the rudists have also received much attention. Moreover, Late Cretaceous Mexican rudist faunas and for having introduced and motivated the rudist faunas are much more diverse than Early and middle Cretaceous authors to follow her with the study of these interesting fossil bivalves. ones. Many papers including descriptions of rudist taxa, most of them new, 2. Material and methods were written during the twentieth century. Some of these descriptions do not fit the modern standards and, although several of the original This research is an exhaustive review of the previously published specimens are preserved and available for study in museums, other papers, including recent ones written by the authors, and observations could not be traced, as well as the stratigraphy of the localities was not on type specimens. The available references and descriptions about always clear or accurate. Taxonomic revisions began to be frequent Mexican Upper Cretaceous rudists in the extant literature were restudied * Corresponding author. Departament de Geologia, Universitat Aurònoma de Barcelona, 08193, Bellaterra, Barcelona, Spain. E-mail addresses: josepmaria.pons@uab.cat (J.M. Pons), pedrogarciab@ciencias.unam.mx (P. García Barrera), aoviedo@uach.mx (A. Oviedo), enric.vicens@uab. cat (E. Vicens). 1 Jose Maria Pons present adress. Apartat de Correus n. 65, 08193, Bellaterra, Barcelona, Spain. jmponsm@gmail.com https://doi.org/10.1016/j.jsames.2021.103237 Received 25 November 2020; Received in revised form 29 December 2020; Accepted 15 February 2021 Available online 26 February 2021 0895-9811/© 2021 The Authors. Published by Elsevier Ltd. This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/). J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 and all the data summarized and organized in a data table (Appendix upper Turonian. It is a relatively abundant fossil in Mexico, where the 1)2. When existing, the repository of type specimens has also been transverse sections of right valves naturally exposed on limestone traced, although not all the collections have already been revisited. The bedding tops are popularly known as ‘patitas de gato’ (cat tracks). Hip­ geologic maps 1:250 000 and 1:50 000 published by the Servicio Geo­ purites mexicana Bárcena (1875) or Hippurites resectus var. mexicana lógico Mexicano are at , , and ) as well as the Léxico Estra­ synonymies. These species have been reported also from Sierra de Bar­ tigráfico de México of the same bosa, Cañada Grande, W Santa Catarina, San Luis-Matehuala, La Calera federal institution. These documents were used to resolve questions (SLP), Cerro La Peña, Cerro Santa Ana (Coah), Cerro de Cadereyta (Qro), about the localities’ original references. Recent regional geological Valle Mezquital (Hgo), Cerro de Escamela (Ver), Apaxco (Mex), and studies were also consulted to confirm or correct both the lithological Yautepec (Mor). The specimens figured by Douvillé (1897, p.230, pl. 32, units reported as a source of the rudist specimens, and their geologic age. Figs. 15, 16) as Hippurites sp. from “near Mexico” correspond to Hip­ Recent papers concerning Mexican Late Cretaceous age-significant purites resectus specimens, probably from Apaxco (Appendix 1). micro- and macrofossils helped for this last purpose. Durania arnaudi (Choffat, 1886-1902) was first described, as Bir­ The institutional repositories of the specimens recognized in this adiolites Arnaudi, from Coz-Juncal, Alcantara, Runa and Monte Serves, study are: FCMP = Museo de Paleontología de la Facultad de Ciencias de Portugal, remarking the differences with Biradiolites cornupastoris Des la UNAM, México; IGM=Colección Nacional de Paleontología at the Moulins, 1826 (=Durania cornupastoris). Besides the type form, the Instituto de Geología de la UNAM, México; LACM = Natural History author distinguished three more varieties (intermedia, Runaensis, Museum of Los Angeles, California, USA; NMNH=Smithsonian, National expansa), indicating the wide morphological variability observed within Museum of Natural History, Washington D.C., USA; MNHN = Muséum the species. The species (within the genus Durania) has been subse­ National d’Histoire Natural, Paris, France; PUAB=Paleontological Col­ quently reported from different upper Cenomanian-Turonian localities lections of the Universitat Autònoma de Barcelona, Bellaterra, Spain; in southern Europe and northern Africa (see Steuber, 2002). Sauvagesia TMM = Bureau of Economical Geology Collection at the Texas Memorial kellyi Jones (1938), from Sierra de la Peña, Coah, is a Durania and a Museum in Austin, Texas, USA; UCBL-EM = Collection de l’École junior synonym of D. arnaudi, as is probably Durania coahuilensis Mül­ Nationale supériéure des Mines de Paris, Université Claude Bernard, lerried (1952). Durania gonzalezi Müllerried (1950b), from El Sotolar, Lyon I, Villeurbanne, France; UMMP=University of Michigan, Museum Manuel Benavides, Chih, as well as other reports of the author as of Paleontology, Ann Harbor, Michigan, USA. D. cornupastoris (Des Moulins) or D. cf. gonzalezi from other Mexican localities (Müllerried, 1950d, 1953) also seem to be junior synonyms of 3. Results D. arnaudi (Appendix 1). Eoradiolites liratus (Conrad, 1852) was proposed, as Hippurites liratus Late Cretaceous rudists species have been reported or described and and figured in external lateral view of the right valve, based on speci­ published from seventeen of the thirty-two Mexican States (Fig. 1). mens from Bhamdun, S of Beyrouth, Lebanon, in rocks attributed to These are: Baja California (BC), Chiapas (Chis), Chihuahua (Chih), lower Turonian. The species, assigned to the genus Radiolites, was Coahuila (Coah), Guerrero (Gro), Hidalgo (Hgo), Jalisco (Jal), México described and figured by Parona (1909) based on both valves of silicified (Mex), Michoacán (Mich), Morelos (Mor), Nuevo León (NL), Oaxaca specimens from Abeih, also S of Beyrouth, a locality considered lower (Oax), Querétaro (Qro), San Luis Potosí (SLP), Tabasco (Tab), Tam­ Cenomanian. The same year, Douvillé (1909) proposed the new genus aulipas (Tamps), and Veracruz (Ver). Rudist families include mainly Eoradiolites, and Douvillé (1910) assigned the species to this genus and Antillocaprinidae, Hippuritidae, Plagioptychidae, and Radiolitidae, but fixed the horizon as being lower Cenomanian. Subsequently, E. liratus also Requieniidae (see: Carter, 2011 for supra-generic classification of has been profusely reported from different localities in Eurasia and the Bivalvia, Mollusca). northern Africa (see Steuber, 2002); its attribution also to the Turonian Cenomanian rudists are not considered in this paper. These rudist in northern Africa corresponds in fact to misidentifications of Eor­ taxa, including mainly Caprinuloideinae among other families, corre­ adiolites schweinfurthi (Zittel, 1883) (Pons et al., 2011). spond to what it is known as “American mid Cretaceous” rudist faunas, The reinterpretation of the radial bands, down-and-outward folds of significantly differing from Late Cretaceous ones. Most Cenomanian the growth lamellae instead of up-and-inward folds, and the outer shell outcrops are often linked to Lower Cretaceous rocks from which, after layer structure of Bauman’s (1958) Radiolites species, newelli and muel­ the published data, are not easy to discriminate. Nevertheless, some lerriedi, from Cerro Corona, Cuernavaca, Mor, indicates that these spe­ considerations on probable Cenomanian radiolitids are included in the cies correspond to the genus Eoradiolites and both show the diagnostic next section 3.1. characters of E. liratus. Radiolites mendozae Bárcena, 1877-1879, cited from different localities, SLP, Qro, Gro, Mich, and Ver, sometimes 3.1. Turonian rudists together with H. resectus, may also be a junior synonym of E. liratus (Appendix 1). Turonian limestones with rudists have been reported in the geolog­ The radiolitids and other rudists described as Turonian by Palmer ical maps of many Mexican States: Chih, Ojinaga Formation; Coah, (1928) from Huescalapa, Jal, are in fact Albian (Alencáster and Gar­ Indidura Formation; Gro, Morelos Formation; Hgo, El Doctor Formation; cía-Barrera, 2008). Jal; Mex, El Doctor Formation; Mich; Mor, Mexcala Formation; Qro, El The requieniids Apricardia mongesi Müllerried, 1947b in Cerro Doctor Formation; SLP, El Abra Formation; and Ver, Guzmantla For­ Peñuela (Ver) and Apricardia sp. in Apaxco (Mex) (Müllerried, 1936b) mation. The authors collected specimens of Hippurites resectus Defrance were reported as occurring together with Hippurites resectus. (1821), together with Durania arnaudi (Choffat, 1886-1902) and, in a Stratigraphy and Geology. The mentioned localities in SLP are mapped lower bed, Eoradiolites liratus (Conrad, 1852), from limestones cropping as El Abra Formation (Albian-Santonian) by SGM (1998c, 1999b, out at Llano las Hormigas, Villa de Guadalupe, SLP (Oviedo, 2006; 2010a). The stratigraphy of mentioned localities in other Mexican states Oviedo et al., 2007). is as yet far to be unambiguously established. The formations: El Doctor Hippurites resectus Defrance (1821) is a cosmopolitan species, origi­ (Albian-Cenomanian), Indidura (Cenomanian-Turonian), Ojinaga (Tur­ nally described from Uchaux, France, considered characteristic of the onian-Santonian), Morelos or Cuautla (Aptian-Cenomanian), Guzmantla (Turonian-Santonian), and Calizas de Escamela (Turonian) have been reported (SGM, 1997b, 1998a, 2000a, 2000b, 2001, 2002), 2010b). 2 References marked with an asterisk (*) are those treated in the Appendix. Hippurites resectus is a helpful Turonian indicator and Durania arnaudi 2 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 Fig. 1. Rudists reported in the Upper Cretaceous of Mexican States. (BC=Baja California; Chih = Chihuahua; Chis = Chiapas; Coah = Coahuila; Gro = Guerrero; Jal = Jalisco; Mex = México; Mich = Michoacán; Mor = Morelos; NL=Nuevo León; Qro = Querétaro; SLP=San Luis Potosí; Tamps = Tamaulipas; Ver = Veracruz) (T = Turonian; S=Santonian; C=Campanian; M = Maastrichtian). occurs in the same bed. Eoradiolites liratus, although reported at some of Ver, assigned by Müllerried (1947b) to Hippurites (Vaccinites) boehmi the same localities, occurs always in lower beds, and may indicate the Douvillé pars., correspond also to V. macgillavry (Appendix 1). Cenomanian. The report of Durania curasavica in the Suchiapa Formation at Tuxtla Gutiérrez, Chis (Alencáster and Michaud, 1987) should correspond to Potosites tristantorresi Alencáster and Pons, in Pons et al. (2010). 3.2. Santonian rudists Stratigraphy and Geology. The locality Sierra de la Ruda, SLP, is mapped as El Abra Formation (Albian-Santonian) by SGM (1999b). The The authors collected specimens of Vaccinites macgillavryi Palmer localities La Esperanza and road from Almolonga to Hueyitlalpan, Gro, (1933) embedded in massive limestones at Sierra de La Ruda, Villa de are mapped as Mexcala Formation (Turonian-Campanian) by SGM Guadalupe, SLP and, in thick thickets from where the specimens can be (1998b). The limestone quarries at Cerro Peñuela, Córdoba, Ver, are easily recovered, at La Esperanza, Zitlala and on the mapped as Guzmantla Formation (Turonian-Santonian) by SGM (2001). Almolonga-Hueyitlalpan road, Mártir de Cuilapan, Gro. A few speci­ mens of Durania curasavica (Martin, 1885) were collected in the last locality (Oviedo, 2006; Oviedo et al., 2007). 3.3. Campanian and Maastrichtian rudists Vaccinites macgillavryi Palmer (1933) was first described from Loma Yucatan, Cuba, poorly described according modern standards, and Campanian and Maastrichtian rudist-bearing rocks have been re­ considered Maastrichtian. Recent research on the stratigraphy of this ported from the states of Baja California (Rosario Formation), Tam­ fossil locality (Rojas et al., 1996; Rojas, 2004) established its age as aulipas (Mendez and San Felipe formations), San Luis Potosí (Cardenas Santonian. Durania curasavica (Martin, 1885) is also reported from the Formation and Temazcal limestones), and Chiapas (Suchiapa, Ocozo­ same locality. coautla, and Angostura formations). Transverse cross-sections of the right valve of V. macgillavry display a The stratigraphy and paleontology of the Cardenas Formation, in wide variability in development and pinching of the two pillars, even SLP, and the Suchiapa, Ocozocoautla, and Angostura formations, in among specimens of the same bed, and the tip of the ligament ridge is Chis, have been intensively investigated during many years until pre­ often difficult to observe because of recrystallization (Oviedo, 2006). sent. The abundant published available data on age-indicative fossils Specimens from different localities in Gro were tentatively assigned (planktic and benthic foraminifers, ammonites, inoceramids) helped to (Alencáster et al., 1987) to European species (V. praegiganteus, date the successive rudist assemblages distinguished within these V. grossouvrei, V. gosaviensis, and V. giganteus), probably because of all formations. these reasons and to the small number of available specimens. On the other hand, the species name macgillavry was erroneously applied (Sirna 3.3.1. Rosario Formation rudists et al., 1990; Pons et al., 1992) to African specimens of V. vesiculosus White (1885) described the new genus and species Coralliochama (Woodward, 1855). The randomly oriented sections on polished slabs in orcutti based on specimens collected by Mr. C.R. Orcutt at Todos los building walls, coming from the quarries at Cerro Peñuela, Córdoba, Santos Bay, BC State, and donated to the NMNH (Appendix 1). The fossil 3 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 locality, Punta Banda, and the morphological variability within the limestones cropping out E of Villa Hidalgo, about 70 km NNE of San Luis species, have been intensively studied (Marincovich, 1975; Aranda Potosí City in SLP State, at Sierras de Álvarez, El Tablón, and Los Librillos, Manteca, 1991; Götz et al., 2005). Coralliochama flouriei Damestoy all of them included in the Sierras and Llanuras Occidentales of the Sierra (1965), from the same locality, is considered a junior synonymy (Mar­ Madre Oriental. The authors collected rudist specimens (Appendix 1) at incovich, 1975). No other rudist species have been reported from this the municipalities of Armadillo de los Infante (Arroyo Tovar in Ranchito locality, nor has this species been reported in Mexico outside BC State. de los Guzmán), Villa Hidalgo (Cerro La Mesa and Hacienda El Durazno in White (1885) indicated the species also occurs near the town of Wallala Paradita del Refugio, El Potrero and Arroyo Temazcal in El Temazcal), (=Gualala), Mendocino County, California. All records are, thus, on the and Guadalcázar (Cerro El Tanque, Cantera de Guadalcázar 1, and Cantera Pacific coast. de Guadalcázar 2 in Charco Blanco, and Arroyo El Aguaje in Aguaje de los Stratigraphy and Geology. The Rosario Formation (Campanian- García) (Oviedo, 2006; Pichardo, 2008; Pons et al., 2010). Maastrichtian) is mapped, besides at Punta Banda, also N and S of Bahia Stratigraphy and Geology. The Temazcal limestones localities are Todos Santos by SGM (2003). The precise stratigraphic location of the mapped as El Abra Formation (Albian-Santonian) by SGM (2000c, rudist beds within the formation is not specifically established. Mar­ 2008b). The early Campanian age attributed to this rudist fauna is due to incovich (1975) indicates a late Campanian to early Maastrichtian age. the presence of Radiolites acutocostata Adkins (1930), because its type locality in Texas is well dated by ammonites indicating the Delawarella 3.3.2. Mendez Formation rudists delawarensis lower Campanian ammonite biozone (Young and Woodruff, Rudists reported from this formation correspond to sparse speci­ 1985) and its absolute age has also been measured, ranging from 83.40 mens, with the collecting site well indicated but without details about its Ma to between 81.71 Ma and 80.54 Ma (Scott, 2005). Potosites tris­ stratigraphic horizon. Stephenson (1922) and Adkins (1930) proposed tantorresi is a new species from that unit, and the age attributed to the several new species of radiolitids or identified other ones previously other species, known from the Caribbean Province, although not so reported from Texas (USA). Their original descriptions and figures are precisely established, does not contradict the early Campanian. A sig­ not sufficient to identify these new species when new specimens from nificant number of species from the Temazcal limestones are repre­ other localities are studied. The specimens (most of the taxa are repre­ sented in the Suchiapa Formation at Chis State (Pons et al., 2016). sented by a single specimen) are kept in fossil collections (NMNH, TMM) available for restudy. A quick visit to the Adkins collection on the 3.3.5. Cardenas Formation rudists occasion of the Seventh International Congress on Rudists in Austin, The stratigraphy and paleontology of the Cardenas Formation, Texas, revealed that Durania aguilae Adkins (1930) is a Chiapasella, thus around Cárdenas town, about 150 km E of San Luis Potosí City, in SLP becoming the senior synonym of Chiapasella trechmanni Mitchell and State, is well known essentially by the research of Böse (1906) and Myers Gunter, 2002, a species described from Jamaica (Mitchell and Pons, (1968). Both authors described many other fossil Mollusca besides 2010). The same may possibly occur with other species: Durania huasteca rudists. Böse compared the stratigraphic succession of the Cardenas beds Adkins (1930) seems to fit within the intraspecific variability of Huasteca with that in Gosau, Austria. Myers defined the Cardenas Formation, ojanchalensis (Myers, 1968), as it is illustrated by Pons et al. (2013). divided it into three informally designated members, and subsequently No more recent reports of rudists from the area have been published recognized three assemblage zones: a lower one (Durania ojanchalensis since then, except Müllerried’s (1931a) Coralliochama n. sp. and Sau­ Zone), a middle one (Arctostrea aguilerae Zone); and an upper one vagesia cf. coloradensis Stephenson. The former was never formalized, (Tampsia floriformis Zone); both lower and upper ones characterized by and the specimen has not been identified in IGM collections (Appendix the presence of rudists. The authors collected specimens in the munici­ 1). palities of Cárdenas (Arroyo la Atarjea, Cárdenas 1, Cárdenas 2, and Stratigraphy and Geology. The Mendez Formation in Tamps, where Cárdenas 3) and Rayón (Amoladeras, Cuchilla Las Palomas, Arroyo Los the specimens described by Stephenson (1922), Adkins (1930), and Terrerillos, and Arroyo el Terrero) (Pons et al., 2013) (Appendix 1). Müllerried (1931a) were collected, is considered Stratigraphy and Geology. The Cardenas Formation is mapped as Campanian-Maastrichtian by SGM (1999a, 2008a). The collecting area Campanian-Maastrichtian by SGM (1997a, 1999a). Pons et al. (2013) is in the Gulf Coastal Plain, where the outcrops of the Mendez Formation dated the two recognized rudist assemblages of the formation respec­ appear at the bottom of gullies, in windows excavated on the Quaternary tively as early and early late Maastrichtian, based on the distribution of alluvial. co-occurring foraminifera (Omaña and Pons, 2000, 2003; Omaña et al., 2009). Some rudist species of the Cardenas Formation were identified in 3.3.3. San Felipe formation rudists the Ocozocoautla (lower Maastrichtian) or the Angostura (upper Two new species of radiolitids, Sauvagesia degolyeri and Durania Maastrichtian) formations in Chis State (Pons et al., 2016, 2017). manuelensis, were proposed respectively by Stanton (1921) and Ste­ phenson (1922) for specimens from the San Felipe Formation in Tamps 3.3.6. Suchiapa Formation rudists (Appendix 1). No more reports of rudists from this formation have been The Upper Cretaceous rudist species from Chiapas State were published since then. Only the type specimens are available for restudy described by Alencáster (1971), upon the numerous specimens collected in museums’ fossil collections (NMNH). Although based only on the along many years by F.K.G. Müllerried and kept, with the corresponding study of original descriptions and figures, Sauvagesia degolyeri Stanton localities indicated, in the IGM collections. Although many rudist taxa (1921) seems to be a senior synonym of Sauvagesia acutocostata Adkins were reported by Müllerried (1931b, c, 1933a, b, 1934, 1936b, 1942, (1930), both corresponding to the genus Radiolites, and Durania man­ 1947a) most of them were not described and/or figured. Alencáster uelensis Stephenson (1922) seems to be a species belonging to the genus (1971) referred to Müllerried’s localities but neither correlating them Chiapasella. nor indicating the corresponding lithological units. Stratigraphy and Geology. Stanton (1921) and Stephenson (1922) The Suchiapa Formation was defined by Michaud (1987) for the indicated the San Felipe Formation, in Tamps, for the fossils they upper part of the limestones attributed to the Sierra Madre Formation, described; nevertheless, the localities they indicated are now mapped as probably with an unconformity between both formations, in the Chiapas Mendez Formation (Campanian-Maastrichtian) by SGM (1999a). The Central Depression, Chis. The authors collected specimens in the outcrops of San Felipe Formation (Coniacian-Santonian) are mapped far Suchiapa Formation at several localities W and N of Tuxtla Gutiérrez, to the W in this map by SGM (1999a). where the Suchiapa Formation is overlain by the Ocozocoautla Forma­ tion: Piedra Parada, SW of Ocuilapa; SW of Emiliano Zapata; Suchiapa 3.3.4. Temazcal limestones rudists river, W of Suchiapa; Santo Domingo river, W of Julián Grajales; S of Temazcal limestones is the informal unit name applied to the Julián Grajales; and Rancho El Comiteco quarry, N of Tuxtla Gutiérrez. 4 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 Also at localities SE of Tuxtla Gutiérrez, where this formation is overlain Berriozábal; and E of Emiliano Zapata, corresponding to the Ocuilapa by the Angostura Formation: road to Paraíso del Grijalva 1, 2, and 3, Platform. Also collections were made at: Cerro Verde, W of Veinte de between Laja Tendida and Paraíso del Grijalva; and road to El Santuario, Noviembre; Colonia José María Morelos y Pavón, SW of Belisario Domí­ SSE of San Francisco Pujiltic (Pons et al., 2016, 2017, 2019) (Appendix nguez Dam; road to Paraíso del Grijalva, between Laja Tendida and 1). Most rudist species from the first group of localities are coincident Paraíso del Grijalva; and SW of San Francisco Pujiltic, corresponding to with those of the Temazcal limestones in SLP, thus early Campanian, the Angostura platform (Pons et al., 2016, 2017). (Appendix 1). while those from the second group are late Campanian. Stratigraphy and Geology. The same rudist species are recorded in Stratigraphy and Geology. Results by Michaud (1987), Michaud and outcrops of both Ocuilapa and Angostura platforms. This indicates that Fourcade (1989), and Cros et al. (1998) are based on a significant the Angostura Formation has the same late Maastrichtian age W and E of number of measured and correlated stratigraphic sections from the Tuxtla Gutiérrez, where it overlies the Ocozocoautla Formation (early Chiapas Central Depression. These authors provided: (1) an accurate Maastrichtian) and directly overlies the Suchiapa Formation (late lithostratigraphic framework of the area (including this formation and Campanian), respectively. Consequently, the Ocozocoautla and the those treated herein in 3.3.7 and 3.3.8); (2) a detailed paleogeographic Angostura formations are not lateral equivalents of each other as pre­ reconstruction, and (3) a reasonable interpretation of the sedimento­ viously maintained (Castro-Mora et al., 1975; Sánchez Montes de Oca, logic evolution of the Chiapas Cretaceous carbonate platform. These 1979; Quezada-Muñetón, 1990; SGM, 2005). The late Maastrichtian age authors proposed that, in the area around Tuxtla Gutiérrez, the original of the Angostura Formation, in both platforms, is also supported by data “mid Cretaceous” carbonate mega-platform (Sierra Madre Formation) on planktic and benthic foraminifers (Omaña and Alencáster, 2019). fragmented into several blocks, each following a different tectono-sedimentary evolution. A drowned block (Tuxtla Gutiérrez 4. Discussion Block) constituted a basin (Tuxtla Gutiérrez Basin) that was filled during Campanian?-Maastrichtian. On the raised block (Angostura Block), and 4.1. Stratigraphy and geology after a long hiatus, an insular shallow carbonate platform (Angostura Platform, Angostura Formation) installed directly above the Sierra The localities with Turonian, Santonian, and early Campanian rud­ Madre Formation. These authors’ tectono-sedimentary model was ists in SLP State are all mapped as El Abra Formation (Albian-Santo­ adopted by Pons et al. (2016) to locate the successive rudist assemblages nian), with the Cardenas Formation (Campanian-Maastrichtian) recognized in the Suchiapa, Ocozocoautla, and Angostura formations in overlying it, by SGM (1998b, 1999b, 2000b, 2000c, 2008, and 2010). the Chiapas Central Depression. The Sierra El Abra, on the easternmost side of the Sierra Madre Oriental, is a carbonate complex in which the El Abra Limestone (carbonate 3.3.7. Ocozocoautla Formation rudists platform) and the Tamabra Limestone (transitional fore-reef to basin The Ocozocoautla Formation crops out N, S, and W of Tuxtla pelagic deposit) were distinguished. Both were first considered as Gutiérrez, Chis. It is mainly composed of siliciclastic rocks, showing members and later as formations. An Albian-Cenomanian age was remarkable facies changes, corresponding to the infilling of the Tuxtla considered for El Abra Formation (Coogan, 1973) although Aguayo Gutiérrez Basin. Proximally, there are conglomerates at the base, fol­ Camargo (1978) reported, on top, a thin and wavy layer containing lowed up section by sandstones, sandy marls with micro-conglomerates planktonic foraminifera dated as late Turonian. intercalations, and a few limestones, with a rudist bed developed in its On the western side of Sierra Madre Oriental, limestones, with suc­ upper part. Distally, the siliciclastic rocks are fine-grained and the marls cessively Turonian, Santonian, and early Campanian rudist faunas, contain inoceramids and ammonites. The authors collected specimens at appear above the limestones with caprinuloideid rudists typical of the El Campo de Tiro and Los Chamaquitos, NW of Ocozocoautla; Abandoned Abra Formation. Oviedo et al. (2007) suggested that successive car­ Quarry, W of Vicente Guerrero; km 178.2, road 145 from Ocozocoautla bonate platforms installed above the El Abra platform, with poorly to Las Choapas; Rancho, NW of Vicente Guerrero; and E of Emiliano developed siliciclastic intervals in between them, until the Cardenas Zapata (Pons et al., 2016, 2017, 2019) (Appendix 1). Formation deposited. Probably, in limited-extent outcrops, these in­ Stratigraphy and Geology. Research on age-indicative fossils in the tervals were interpreted as Cardenas Formation. As no other formation: planktic and benthic foraminifers (Omaña and Pons, 2003; well-defined lithological units were available in the region, Pons et al. Omaña, 2006); ammonites (Pons et al., 2016); and inoceramid bivalves (2010) adopted the informal Temazcal limestones for those with early (Alencáster and Omaña, 2006), indicates that the age of the Ocozo­ Campanian rudists. No other lithological units have been used for the coautla Formation rudist bed is early Maastrichtian. upper Turonian or the Santonian rudist limestones in the area. The Indidura Formation and the San Felipe Formation are recognized for 3.3.8. Angostura Formation rudists limestones, respectively at the Mesozoic Basin of Central Mexico and at The Angostura Formation is mainly composed of shallow-water the eastern margin of the Sierra Madre Oriental. Omaña et al. (2013) in a limestones and crops out in the N 55◦ W trending synclines of the study of the Cenomanian/Turonian interval foraminifers W of the Chiapas Central Depression, Chis. At the central part of the Ocozo­ Valles-San Luis Potosi Platform, distinguished the El Abra Formation coautla Syncline, the Angostura Formation overlies the Ocozocoautla (Albian-Cenomanian), the Soyatal Formation (Turonian-Santonian), and Formation and interdigitates with the sandy bioclastic limestones of the the Cardenas Formation (Campanian-Maastrichtian). The Cardenas Juan Crispin Formation. It is interpreted as a marginal shallow car­ Formation is considered early-late Maastrichtian in Cárdenas (Pons bonate platform (Ocuilapa Platform) surrounded by a belt of cross- et al., 2013). Definitively, the stratigraphy and sedimentology of the bedded sandy bioclastic limestones prograding basinwards. At both Turonian-Campanian rocks in this area is not satisfactorily known. flanks of the Grijalva Syncline, the Angostura Formation overlies un­ No major problems arise with the rest of Campanian and Maas­ conformably the Suchiapa Formation, with bauxitic laterites developed trichtian formations, only the lack of a precise stratigraphic position of between them both. It is interpreted as an insular shallow carbonate the rudist beds within some long-range units, as the Rosario and Mendez platform (Angostura Platform) (Michaud, 1987; Michaud and Fourcade, formations (Campanian-Maastrichtian) and the San Felipe Formation 1989; Pons et al., 2016). Rudists are abundant in the limestones of both (Coniacian-Santonian) (SGM, 1999a; 2003a). The Cardenas Formation, platforms. with the biostratigraphy and depositional history by Myers (1968) and The authors collected specimens at: Cerro Testigo, Old Quarry, Rancho the taxonomic revision of the rudists by Pons et al. (2013), is reasonably La Peregrina, and Búfalo de Piedra, respectively at km 195.5, 195.3, 189, clear. Similarly, the Suchiapa, Ocozocoautla, and Angostura formations, and 187.4, road 145 from Ocozocoautla to Las Choapas; W of Vicente with the lithostratigraphic framework, paleogeographic reconstruction, Guerrero; Quarries, W of Berriozábal; Quarries El Tesoro, NW of and sedimentologic evolution of the Chiapas Carbonate Platform by 5 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 Michaud and Fourcade (1989), and the successive rudist assemblages Table 1 distinguished by Pons et al. (2016), are also clear. Rudist species recognized in the Mexican Upper Cretaceous (Turonian- Maastrichtian). 4.2. Taxonomy Species Formation Localities State Age ANTILLOCAPRINIDAE Four rudist families: Antillocaprinidae MacGillavry, 1937, Hippur­ Alencasteria Suchiapa Laja Tendida to Chis lC itidae Gray, 1848, Plagioptychidae Douvillé, 1888, and Radiolitidae macrotubularis Paraíso del d’Orbigny (1842), are well represented and diverse in the Mexican Mitchell (2013) Grijalva Upper Cretaceous. The knowledge on the species of each family is not Antillocaprina Angostura Ocozocuautla to Chis lM uniform and many species need a restudy. Family Requieniidae Kutassy, quadrangularis ( Las Choapas Whitfield, 1897a) 1934 is rarely reported. Antillocaprina Angostura Ocozocuautla to Chis lM suboccidentalis Las Choapas 4.2.1. Antillocaprinidae (Table 1) Chubb (1967) The modern study of the Mexican Upper Cretaceous Anti­ Caenosarcolites Angostura Ocuilapa Chis lM llocaprinidae is still to be undertaken. Only the few specimens described oddsensis ( Stephenson, by Alencáster (1971) and those by García Barrera and Avendaño in 1938) García Barrera et al. (1998) have been considered in the revision of the Oligosarcolites Suchiapa Laja Tendida to Chis lC Antillocaprinidae by Mitchell (2013). Modern papers on Mexican rudists monotubularis Paraíso del paleontology (Oviedo, 2006; Pons et al., 2010, 2016) include abundant Mitchell and Grijalva Gunter (2006) references to Antillocaprinidae genus indeterminate, Antillocaprinidae ? Parasarcolites Angostura Rancho La Chis lM indeterminate, Antillocaprina sp., Titanosarcolites sp., and some uncer­ adkinsoni Peregrina tain specific determinations, from different Campanian and Maas­ Mitchell and trichtian formations. Also, many undescribed Antillocaprinidae Gunter (2006) specimens are kept in the collections of Mexican museums. All this in­ Rotacaprina? Angostura Ocozocoautla to Chis lM trilobata (García Las Choapas dicates that Mexican Antillocaprinidae may be diverse but are insuffi­ Barrera and ciently known. It is time to start this study; the general revision by Avendaño in Mitchell (2013) may be used as a base. García-Barrera The genus Alencasteria Mitchell, 2013 (type species Alencasteria et al., 1998) Stellacaprina Suchiapa Laja Tendida to Chis lC macrotubularis Mitchell, 2013) was established based on Mexican ma­ macgilavryi ( Paraíso del terial with part of the specimens from Vega del Paso, Chis (Suchiapa Alencáster, 1971) Grijalva Formation) previously figured as Titanosarcolites macgillavryi by Titanosarcolites Angostura Ocozocoautla to Chis lM Alencáster (1971, pl. 5, Figs. 1, 4, pl. 18, Fig. 4, pl. 19, Fig. 1). The other giganteus Las Choapas specimens (Alencáster, 1971, pl. 4, Fig. 4, pl. 5, Figs. 2–3) were tras­ Whitfield (1897a) Antillocaprinidae Temazcal lst. Temazcal SLP eC ferred to genus Stellacaprina Mitchell 2013 as Stellacaprina macgillavryi genus (Alencáster, 1971) or (Alencáster, 1971, pl. 5, Fig. 5) assigned to Oli­ indeterminate [in gosarcolites monotubularis (Mitchell and Gunter, 2006). Another Anti­ Pons et al., 2010] llocaprinidae specimen described although not figured by Alencáster Antillocaprinidae Suchiapa El Comiteco Chis lC indet. [in Pons Quarry (1971) was assigned by Mitchell (2013) to Caenosarcolites oddsensis et al., 2016] (Stephenson, 1938). “Postcaprinula” Suchiapa Laja Tendida to Chis lC The species Antillocaprina trilobata García Barrera and Avendaño, in Alencáster [in Paraíso del García Barrera et al, 1998 from the Angostura Formation (Chis) was in collection IGM] Grijalva part (García-Barrera et al., 1998; text-fig. 3.1, 2, 4) transferred to genus HIPPURITIDAE Rotocaprina? and in part (García-Barrera et al., 1998; text-fig. 3.3, 3.5) Barrettia gigas Suchiapa Laja Tendida to Chis lC assigned to Antillocaprina suboccidentalis Chubb (1967) by Mitchell Chubb (1955) Paraíso del (2013). Grijalva The species Antillocaprina pugniformis (Palmer, 1933), considered a Barrettia monilifera Suchiapa Laja Tendida to Chis lC junior synonymy of Antillocaprina quadrangularis (Whitfield, 1897a) by Woodward Paraíso del (1862) Grijalva Mitchell (2013), was described from the Angostura Formation (Chis) by Barrettia ruseae Temazcal Guadalcázar; El SLP; eC Oviedo (2006). Recently collected specimens from the Angostura For­ Chubb (1967) limestone; Comiteco Quarry Chis mation are herein tentatively assigned to ?Parasarcolites adkinsoni Suchiapa. Mitchel and Gunter, 2006. Specimens from the Suchiapa Formation at Caribbea Cardenas Cárdenas SLP lM muellerriedi ( Vega del Paso in the IGM collection labelled as “Postcaprinula” by Vermunt, 1937) Alencáster and new material recently collected has not been described Hippurites resectus El Abra, Llano las SLP; T up to now. Defrance (1821) Cuautla; El Hormigas, Sierra Coah; Doctor; de Barbosa, Qro; 4.2.2. Hippuritidae (Table 1) Morelos; Cañada Grande, W Mex; Guzmantla Santa Catarina; Mor; Only one species, the cosmopolite Turonian Hippurites resectus Cerro de Ver Defrance (1821) is common with the Eurasian Hippuritidae, and it is one Cadereyta; of the more widespread hippuritid species in Mexico (Coah; Hgo; ?Jal; Apaxco; Yautepec; Mex; Mor; Qro; SLP). Genus Vaccinites is also common with Eurasia but Cerro Escamela Parastroma Suchiapa Laja Tendida to Chis lC the two species, V. macgillavryi Palmer (1933) and V. vermunti MacGil­ trechmanni Chubb Paraíso del lavry, 1937 (both SLP and Chis; the former also Ver), are only known (1967) Grijalva from America (Caribbean Province). Some other reports of Eurasian Praebarrettia armini Cardenas Rayón SLP eM hippuritid species in Mexico (Hippurites cf. incisus Douvillé, Vaccinites (Götz and giganteus major Toucas, and Vaccinites boehmi Douvillé) by Müllerried Mitchell, 2009) (1930b, 1933b, 1936b, 1947b) are misidentifications. (continued on next page) 6 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 Table 1 (continued ) Table 1 (continued ) Species Formation Localities State Age Species Formation Localities State Age Praebarrettia Ocozocoautla Ocozocoautla Chis eM Los Hornos; Sierra sparcilirata de la Peña, Sierra Whitfield (1897b) de Santa Ana Torreites sanchezi ( Temazcal Armadillo de los SLP eC Durania curasavica Mexcala Almolonga- Gro S Douvillé, 1927) limestone Infante, Martin (1885) Hueyitlalpan Guadalcázar Durania huasteca Mendez; Hacienda Tamps M Vaccinites El Abra; Sierra de La Ruda; SLP; S Adkins (1930) Angostura Cacalilao macgillavryi Mexcala; La Esperanza, Gro; Ver Durania manuelensis San Felipe Manuel Station Tamps C Palmer (1933) Guzmantla Almolonga- Stephenson Hueyitlalpan; (1922) Cerro Peñuela Eoradiolites liratus ( El Abra; El Villa de SLP; Cen?- Vaccinites vermunti Temazcal Armadillo de los SLP; eC Conrad, 1852) Doctor; Morelos Guadalupe Qro; T MacGillavry, limestone; Infante, Villa Chis Gro; 1937 Suchiapa Hidalgo, Mich; Guadalcázar; El Ver; Jal Comiteco Quarry Huasteca Cardenas; Cárdenas, Rayón; SLP; eM ojanchalensis ( Ocozocoautla Ocozocoautla Chis PLAGIOPTYCHIDAE Myers, 1968) Coralliochama Cardenas Cárdenas SLP M Potosites Temazcal lst.; Armadillo de los SLP; eC gboehmi Böse tristantorresi Suchiapa Infante, Villa Chis (1906) Alencáster and Hidalgo, Coralliochama Rosario Punta Banda, BC lC- Pons, in Pons Guadalcázar; El orcutti White Bahia Todos eM et al., 2010 Comiteco Quarry (1885) Santos Radiolites Temazcal lst.; Armadillo de los SLP; eC Mitrocaprina Cardenas; Cárdenas, Rayón; SLP; lM acutocostata ( Suchiapa Infante, Villa Chis tschoppi (Palmer, Angostura Old Quarry, Chis Adkins, 1930) Hidalgo, 1933) Ocozocoautla to Guadalcázar; El Las Choapas Comiteco Quarry Plagioptychus Suchiapa Laja Tendida to Chis lC “Radiolites” Angostura Colonia José Chis lM antillarum ( Paraíso del macroplicatus María Morelos y Douvillé, 1927) Grijalva Whitfield (1897a) Pavón Plagioptychus fragilis Angostura Rancho La Chis lM Sauvagesia belti Mendez Chocoy Station Tamps M Chubb (1967) Peregrina Stephenson Plagioptychus Ocozocoautla Ocozocoautla Chis eM (1922) muellerriedi Sauvagesia Mendez Hacienda Chocoy Tamps M Alencáster (1971) coloradensis Mitrocaprina sp. [in Suchiapa Laja Tendida to Chis lC Stephenson Pons et al., 2016] Paraíso del (1922) Grijalva Sauvagesia degolyeri San Felipe Haciendas Tamps C Plagioptychus sp. [in Temazcal Villa Hidalgo SLP eC Stanton (1921) Chocoy, Las Flores Pons et al., 2010] limestone Sauvagesia morgani Mendez Hacienda Las Tamps M Plagioptychidae Temazcal Villa Hidalgo SLP eC Adkins (1930) Flores genus limestone Tampsia bishopi Mendez Hacienda Las Tamps M indeterminate [in Stephenson Flores Pons et al., 2010] (1922) Tampsia chocoyensis Mendez Hacienda Chocoy Tamps M RADIOLITIDAE Stephenson Alencasterites Suchiapa Laja Tendida to Chis lC (1922) mooretownensis ( Paraíso del Tampsia floriformis Cardenas Cárdenas SLP lM Trechmann, Grijalva Myers (1968) 1924) Thyrastylon Angostura Ocozocoautla to Chis lM Biradiolites aguilerae Cardenas Cárdenas SLP M adhaerens Las Choapas Böse (1906) Whitfield (1897a) Biradiolites Cardenas; Cárdenas, Rayón; SLP; eM- “Thyrastylon” Suchiapa; Laja Tendida to Chis lC- cardenasensis Ocozocoautla; Ocozocoautla; Chis lM nicholasi Ocozocoautla Paraíso del eM Böse (1906) Angostura Ocozocoautla to Whitfield (1897a) Grijalva, Rio Las Choapas Suchiapa; Bournonia baileyi Suchiapa Laja Tendida to Chis lC Ocozocoautla Chubb,1967 Paraíso del Trechmannites Cardenas; Cárdenas, Rayón; SLP; eM- Grijalva rudissimus ( Ocozocoautla Ocozocoautla Chis lM Bournonia cancellata Angostura Ocozocoautla to Chis lC Trechmann, Whitfield (1897a) Las Choapas 1924) Chiapasella aguilae ( Mendez; San Antonio Tamps; M REQUIENIIDAE Adkins, 1930) Angostura Rayón; Cerro Chis Verde Apricardia mongesi Guzmantla Cerro Peñuela, Ver T Chiapasella Ocozocoautla Ocozocoautla Chis eM- Müllerried Córdoba radiolitiformis ( lM (1947b) Trechmann, Apricardia sp. [in El Doctor Apaxco Mex T 1924) Müllerried, Chiapasella sp. [in Suchiapa Piedra Parada, Chis eC 1936b] Pons et al., 2016] Suchiapa Durania arnaudi El Abra; Llano las SLP; T Age abbreviations: Cen = Cenomanian; T = Turonian; S=Santonian; Choffat Ojinaga; Hormigas; El Chih; C=Campanian; M = Maastrichtian; e = early; l = lateMexican States abbrevia­ (1886-1902) Morelos; Sotolar; Mor; tions: BC=Baja California; Chih = Chihuahua; Chis = Chiapas; Coah = Coahuila; Indidura Tlaquiltenango, Coah Gro = Guerrero; Jal = Jalisco; Mex = México; Mich = Michoacan; Mor = Morelos; NL=Nuevo León; Qro = Querétaro; SLP=San Luis Potosí; Tamps = Tamaulipas; Ver = Veracruz. 7 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 Three multiple-fold hippuritid genera, characteristic of the Carib­ cited. Nevertheless, some of the species assigned to these genera have bean Province, are represented in Mexico: Barrettia, Parastroma, and been subsequently transferred to other ones (Mitchell and Pons, 2010). Praebarrettia. The former with one species, B. ruseae Chubb (1967), in Probably, a careful revision of the original material, single specimens the lower Campanian (Temazcal limestones, SLP, and Suchiapa For­ insufficiently described and figured, may lead to further changes. mation, Chis) and two more, B. gigas Chubb (1955) and B. monilifera The only species of the genus Eoradiolites Douvillé, 1909 reported Woodward (1862), in the upper Campanian occurring together with from the Mexican Upper Cretaceous, Eoradiolites liratus (Conrad, 1852), Parastroma trechmanni Chubb (1967) (Suchiapa Formation, Chis). occurs in the Cenomanian? or Turonian? of different localities (SLP, Qro, Praebarrettia armini (Götz and Mitchell, 2009) and P. sparcilirata Mor, Gro, Mich, and Ver) and is the senior synonymy of Mexican species (Whitfield, 1897b) both occur in the lower Maastrichtian, but in originally assigned to genus Radiolites: R. mendozae Bárcena, 1877-1879; different formations and areas (Cardenas Formation, SLP, the former R. newelli and R. muellerriedi Bauman 1858. and Ocozocoautla Formation, Chis, the latter). Reports of Biradiolites or Distefanella, lombricalis d’Orbigny (1847) or, Torreites sanchezi (Douvillé, 1927) from the lower Campanian lumbricalis Parona, 1901, sometimes with exchanged genera and species Temazcal limestones, SLP, and Caribbea muellerriedi (Vermunt, 1937) names, in Oax and Chis (Böse, 1910; Müllerried, 1932b, 1933b, 1936b, from the upper Maastrichtian Cardenas Formation, SLP, are peculiar 1950a; Chubb, 1959), probably correspond to Alencasterites moor­ hippuritids. The former by lacking the pore-canal system of the outer etownensis (Trechmann, 1924). Some other reports of Eurasian radiolitid shell layer, and having canals in the inner shell layer, of the left valve. species in Mexico (Sauvagesia cf. da Rio Catullo, Radiolites aff. lusitanicus The latter by having non-compact structure in the outer shell layer of the Bayle, Biradiolites canaliculatus d’Orb., Radiolites cf. subradiosus Toucas, right valve. Radiolites cf. galloprovincialis Matheron, and Praeradiolites aff. sub­ Hippurites llamasi Müllerried (1948a), from Valle del Mezquital, Hgo, coquandi Toucas) by Müllerried (1936b), only corresponding to names seems to be an indeterminate radiolitid. without description or figure, are also misidentifications. Müllerried’s (1946, 1947b) new species Neoradiolites ordoñezi, 4.2.3. Plagioptychidae (Table 1) Bournonia carrilloi, and Durania mexicana are based on sections in The genus Coralliochama White, 1885 (type species Coralliochama building slabs, undeterminable because not showing sufficient diag­ orcutti) was established based on specimens from the Rosario Formation nostic characters. at Punta Banda, Bahia de Todos los Santos, BC. Böse (1906) established Several American species of the genus Biradiolites d’Orbigny Coralliochama gboehmi based on specimens from the Cardenas Formation (1847-51) have been reported, and accurately described, from the at Cárdenas. Müllerried (1931a) described as Coralliochama n. sp. two Maastrichtian Cardenas Formation, SLP, the upper Campanian Suchiapa fragments of right valves from Rayón (=San Antonio Rayón), Tamps. Formation, Chis, the lower Maastrichtian Ocozocoautla Formation, Chis, The author repeatedly cited the species (Müllerried, 1932a, 1947a) but and the upper Maastrichtian Angostura Formation, Chis. Similarly, for never denominate it. Probably it coincides with Adkins’ (1928) report of species of genus Bournonia Fischer, 1887, from the upper Campanian Coralliochama cf. boehmi from the Mendez Formation, Tamps. Cor­ Suchiapa Formation, Chis, and the upper Maastrichtian Angostura For­ alliochama is an exclusively American genus. mation, Chis. Species of genera Mitrocaprina and Plagioptychus are widespread in Five radiolitid genera are based on Mexican specimens: (1) Tampsia Eurasia but none of them occurs in America, although some Stephenson, 1922, from the Maastrichtian Mendez Formation in Tamps; (P. toucasianus, P. arnaudi) were erroneously reported (Müllerried, (2) Chiapasella Müllerried, 1931b, from the lower Maastrichtian Oco­ 1933a, b, 1936b, 1947a; Chubb, 1959). Three Plagioptychus species are zocoautla Formation in Chis; (3) Huasteca Pons et al., 2013, from the known from Mexico: (1) Plagioptychus antillarum (Douvillé, 1927) from lower Maastrichtian Cardenas Formation in SLP; (4) Potosites Alencáster the upper Campanian Suchiapa Formation, (2) Plagioptychus muellerriedi and Pons in Pons et al. (2010), from the lower Campanian Temazcal Alencáster (1971) from the lower Maastrichtian Ocozocoautla Forma­ limestones in SLP; and (5) Trechmannites Pons et al., 2013 from the tion, and (3) Plagioptychus fragilis Chubb (1967) from the upper Maas­ Maastrichtian Cardenas Formation in SLP. None of them occur outside trichtian Angostura Formation, all in Chis. Mitrocaprina tschoppi America. Two more genera are also exclusively American: Alencasterites (Palmer, 1933) occurs in the Maastrichtian Cardenas Formation, SLP, Pons et al., 2016, reported from the upper Campanian Suchiapa For­ and Angostura Formation, Chis. Several other taxa have been reported in mation, Chis; and Thyrastylon Chubb, 1956 from the upper Maas­ open nomenclature: Mitrocaprina sp. from the lower Campanian trichtian Angostura Formation, Chis. The species originally assigned to Temazcal limestones, in SLP, and Suchiapa Formation, in Chis; Pla­ Radiolites (Lapeirousia) nicholasi by Whitfield (1897a) and also attributed gioptychus sp. and Plagioptychidae genus indeterminate from the to Macgillavryia Rojas et al., 1996, reported from the upper Campanian Temazcal limestones, SLP (Pons et al., 2010, 2016). Suchiapa Formation and the lower Maastrichtian Ocozocoautla Forma­ The Mexican Plagioptychidae species already named are fully tion, Chis, is tentatively assigned to Thyrastylon? nicholasi (Whitfield, described and illustrated, according to modern standards (Marincovich, 1897a). 1975; Pons et al., 2013, 2017). Thus, any new find of them elsewhere Besides Chiapasella radiolitiformis (Trechmann, 1924) from the lower may be unambiguously identified. Nevertheless, the abundant open Maastrichtian Ocozocoautla Formation, and Chiapasella aguilae (Adkins, nomenclature reports could indicate that new taxa are waiting to be 1930) from the upper Maastrichtian Angostura Formation, another, still described. unnamed, Chiapasella occurs in the early Campanian Suchiapa Forma­ tion, all of them in Chis (Pons et al., 2016). Müllerried (1931b) proposed 4.2.4. Radiolitidae (Table 1) the species pauciplicata, for a specimen from an unknown horizon at The knowledge of the abundant Mexican Upper Cretaceous Radio­ Cerro de la Silla, Monterrey, NL; the specimen, not figured and described litidae reported during last century is, up to now, unsatisfactory. Most as a rock fragment with three embedded incomplete specimens, has not original descriptions and illustrations do not fit modern standards, not been found in the collections of the IGM. allowing their unambiguous identification. Only some of the specimens are available in Museum collections for restudy, and some of the pro­ 4.2.5. Requieniidae (Table 1) posed new species were never formalized. Thus, the results herein, The Requieniidae are rarely reported from the Mexican Upper mainly based on the original publications, represent only tentative re- Cretaceous, and poorly described. Apricardia mongesi Müllerried assignations or synonymies. (1947b), has been based on randomly oriented sections in polished slabs Among the genera being common in Eurasia, Radiolites Lamarck, of building walls, coming from quarries of the Guzmantla Formation at 1801 has been rarely mentioned in the Upper Cretaceous of Mexico, Cerro Peñuela, Ver. Apricardia sp. was reported by Müllerried (1936b) while Sauvagesia Bayle 1857 and Durania Douvillé, 1908 are the most from Apaxco, Mex, a locality with Hippurites resectus. 8 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 4.3. Perspectives the two stages or to the lower or upper part of them. They have been reported mainly in BC, SLP, Tamps, and Chis states, and also in neigh­ The study, mainly field work, of the Turonian, Santonian, and lower boring areas of NL, Oax, and Tab states. Campanian limestones with rudists, above the mid Cretaceous El Abra Research on the rudist species and its consequent record helped to Formation, in SLP State should be reprised with a modern scope. The improve the stratigraphy in some areas and to point out or clear up present knowledge of the rudists may help. several age issues in other, thus contributing to a better understanding of Also, more field work is necessary to clear up the stratigraphy of Mexican geology. rudist beds in Sierra de la Peña and Sierra Santa Ana, Coah, and Cerro A lower and an upper Maastrichtian interval have been distinguished Peñuela, Ver. in the Cardenas Formation at SLP. The relationship of Coralliochama orcutti White, 1885 with other Lower Campanian, upper Campanian, lower Maastrichtian, and Coralliochama species should be investigated. A better knowledge of the upper Maastrichtian intervals have been distinguished in what was stratigraphic distribution of all them may be helpful. previously considered Campanian-Maastrichtian in Chis. Concerning the Mendez Formation in Tamps, both Stephenson The relationship between the Ocozocoautla and Angostura forma­ (1922) and Stanton (1921) collections, at NMNH, and Adkins (1930) tions in Chis, previously considered a lateral facies change, has been collection at TMM, need imperatively a careful restudy. Several of their evidenced, the Angostura Formation (upper Maastrichtian) laying above species have only rarely (if any) been used subsequently, and a quick the Ocozocoautla Formation (lower Maastrichtian). look to some of the specimens or an attentive examination of their fig­ The record of Turonian, Santonian, and lower Campanian rudist ures suggest the presence of several synonymies or erroneous generic assemblages in what is mapped as El Abra Formation (Albian-Santonian) ascriptions. Although it is desirable, not much should be expected from in SLP evidenced that the stratigraphy and the sedimentological struc­ new collecting in the area, as far as the original material mostly corre­ ture of this Formation needs a different explanation. sponds to occasional find of single specimens. Also, the difficulties in assigning a lithological unit to the rudist Being the stratigraphy of the rudist assemblages in Chiapas species records indicates that, probably, other formations used in the (Suchiapa, Ocozocoautla, and Angostura formations) reasonably estab­ Upper Cretaceous (or the mid Cretaceous) of the western margin of the lished within a consistent sedimentological framework (Pons et al., ‘Valles-San Luis Potosi Platform’, as the San Felipe and Indidura for­ 2016), the subsequent research should focus on improving the taxonomy mations, need to be re-evaluated. of the rudists. Only a limited number of Chiapas specimens were included in Mitchell’s (2013) revision of the Antillocaprinidae. Pla­ Author statement gioptychidae and Hippuritidae have been recently revised (Pons et al., 2017, 2019), but the Radiolitidae, the larger family, is waiting for a deep Jose Maria Pons, have contributed on all the aspects and phases of restudy providing complete descriptions and accurate figures. the research, ending with the elaboration of text, figure, and tables, and subsequent revision, coordinated by J.M.Pons. Pedro García-Barrera, 5. Conclusion have contributed on all the aspects and phases of the research, ending with the elaboration of text, figure, and tables, and subsequent revision, Around sixty rudist species are recognized up today in the Mexican coordinated by J.M.Pons. Angélica Oviedo, have contributed on all the Upper Cretaceous (Table 1): Antillocaprinidae (12); Hippuritidae (11); aspects and phases of the research, ending with the elaboration of text, Plagioptychidae (9); Radiolitidae (27); and Requieniidae (2). A few of figure, and tables, and subsequent revision, coordinated by J.M.Pons. them are in open nomenclature although supposed different from the Enric Vicens, have contributed on all the aspects and phases of the others. Several synonymies are evidenced among previously proposed research, ending with the elaboration of text, figure, and tables, and species (Appendix 1). subsequent revision, coordinated by J.M.Pons. Only three species are common with Eurasia: Eoradiolites liratus (Conrad, 1852), Hippurites resectus Defrance (1821) and Durania arnaudi (Choffat, 1886-1902). The first probably in the Cenomanian and the Declaration of competing interest other in the Turonian, are widespread in outcrops of several Mexican States. The authors declare that they have no known competing financial No coniacian rudists have been reported in Mexico interests or personal relationships that could have appeared to influence The only two Santonian species, Vaccinites macgillavryi Palmer the work reported in this paper. (1933) and Durania curasavica (Martin, 1885) recognized in SLP and Gro states, are American species of cosmopolitan genera. Acknowledgments All the Campanian and Maastrichtian rudists are exclusively Amer­ ican species, either ascribed to Eurasian (9) or to American (21) genera. This research did not receive any specific grant from funding The four rudist families represented are remarkably diversified: Anti­ agencies in the public, commercial, or not-for-profit sectors.J. Gallemí llocaprinidae (9 genera, 12 species); Hippuritidae (5 genera, 9 species); assisted with the first English version of the text. Concerns and sugges­ Plagioptychidae (4 genera, 9 species); Radiolitidae (11 genera, 24 spe­ tions by the reviewers PWS and RC and the Regional Editor M. Martini, cies). The stratigraphic range of most of the species is limited to one of helped to improve the manuscript and are gratefully acknowledged. Appendix 1. Published papers dealing with Mexican Upper Cretaceous rudists Author/Localities Reported taxa Current identification Repository Bárcena (1875) Mex, Apaxco; Mor, Yautepec; Qro; Hgo; SLP, Hippurites mexicana n. sp. Hippurites resectus Defrance (1821) Sierra de Barbosa, Cañada Grande Qro, Cerro de Cadereyta Hippurites calamitiformis n. sp. Hippurites resectus Defrance (1821) [non Monopleura calamitiformis (Bárcena) from Coalcoman, Mich] (continued on next page) 9 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Bárcena (1877-1879) Mex, Apaxco; Mor. Yautepec; Qro; Hgo; SLP, Hippurites mexicana Barcena Hippurites resectus Defrance (1821) Sierra de Barbosa, Cañada Grande SLP, Sierra de Barbosa; Qro; Gro; Mich; Ver Radiolites Mendozae n. sp. ?Eoradiolites liratus (Conrad, 1852) (sometimes together with Hippurites) Qtro, Cerro de Cadereyta Hippurites calamitiformis n. sp. Hippurites resectus Defrance (1821) [non Monopleura calamitiformis (Bárcena) from Coalcoman, Mich] White (1885) NMNH BC, Bahia Todos Santos Coralliochama orcutti n. sp. Coralliochama orcutti White (1885) 186661–186665 Douvillé (1891-1897) UCBL-EM Mex, near Mexico [?Apaxco] Hippuritessp. Hippurites resectus Defrance (1821) Böse (1899) Ver, calizas de Escamela Hippurites (Vaccinites) mexicanus Barc. Hippurites resectus Defrance (1821) Hippurites sp. Böse (1906) ? +IGM SLP, Cardenas Formation Coralliochama G. Boehmi n. sp. Coralliochama gboehmi Böse (1906) 1350,1351,1974 Radiolites Austinensis Roem. ? Biradiolites Aguilerae n. sp. Biradiolites aguilerae Böse (1906) 1354,1355,1357 Biradiolites Cardenasensis n. sp. Biradiolites cardenasensis Böse (1906) 1976,1977 Biradiolites Potosianus n. sp. Biradiolites cardenasensis Böse (1906) 1978 Böse (1910) Oax, Ejutla District Biradiolites lumbricalis d’Orb. ?Alencasterites mooretownensis (Trechmann, 1924) Stanton (1921) NMNH Tamps, San Felipe Formation Sauvagesia degolyeri n.sp. ?Radiolites degolyeri (Stanton, 1921) 32482 Stephenson (1922) NMNH Tamps, Mendez shale Tampsia bishopi n. gen, n. sp. Tampsia bishopi Stephenson (1922) 32499 Tampsia chocoyensis n. sp. Tampsia chocoyensis Stephenson (1922) 32500,32501 Sauvagesia belti n. sp. Sauvagesia belti Stephenson (1922) 32504 Sauvagesia coloradensis n. sp. Sauvagesia coloradensis Stephenson (1922) 32505 Tamps, San Felipe Formation Sauvagesia degolyeri Stanton (?) ?Radiolites degolyeri (Stephenson, 1922) Durania manuelensis n. sp. ?Chiapasella manuelensis (Stephenson, 1922) 32506 Adkins (1928) Tamps, Mendez Formation ?Durania austinensis (Römer) ? Durania manuelensis Stephenson ?Chiapasella manuelensis (Stephenson, 1922) Sauvagesia degolyeri Stanton ?Radiolites degolyeri (Stanton, 1921) Sauvagesia belti Stephenson Sauvagesia belti Stephenson (1922) Sauvagesia coloradensis Stephenson Sauvagesia coloradensis Stephenson (1922) Tampsia bishopi Stephenson Tampsia bishopi Stephenson (1922) Coralliochama cfr. boehmi Böse Coralliochama cf. Coralliochama gboehmi Böse (1906) Adkins (1930) TMM Tamps, Mendez Formation Durania aguilae n. sp. Chiapasella aguilae (Adkins, 1930) BEG2066.5 Durania huasteca n. sp. ?Huasteca huasteca (Adkins, 1930) BEG Sauvagesia morgani n. sp. Sauvagesia morgani Adkins (1930) BEG Müllerried (1930a) Mor, Yautepec; Mex, Apaxco Hippurites (Hippuritella H. Douvillé; Orbignya Hippurites resectus Defrance (1821) Toucas) resectus nov. var. mexicana (Bárcena) Müllerried (1930b) SLP, Cárdenas Hippurites (Hippuritella H. Douvillé; Orbignya ?Caribbea muellerriedi (Vermunt, 1937) Toucas) cf. incisus (H. Douvillé) Toucas Müllerried (1930c) Mex, Apaxco Hippurites (Hippuritella H. Douvillé; Orbignya Hippurites resectus Defrance (1821) Toucas) calamitiformis Bárcena Müllerried (1931a) Tamps, Rayón [ = San Antonio Rayón] Coralliochama n. sp. ?Coralliochama sp. Sauvagesia cf. coloradensis Stephenson Sauvagesia cf. coloradensis Stephenson, 1922 Müllerried (1931b) Chis, Ocozocoautla Chiapasella n. gen. radiolitiformis Chiapasella radiolitiformis (Trechmann, 1924) (Trechmann) Chiapasella n. sp.? ?Chiapasella radiolitiformis (Trechmann, 1924) NL, Monterrey, Cerro de la Silla Chiapasella pauciplicata n. sp. ?Chiapasella pauciplicata Müllerried (1931b) Müllerried (1931c) Chis, Ocozocoautla Pseudobarrettia chiapasensis n. gen., n. sp. ?Praebarrettia sparcilirata (Whitfield, 1897b) Müllerried (1932a) BC, Coralliochama orcutti (White, 1885) Coralliochama orcutti White (1885) SLP, Cárdenas Coralliochama G. Boehmi Boese, 1906 Coralliochama gboehmi Böse (1906) Tamps, Rayón [ = San Antonio Rayón] Coralliochama n. sp. ?Coralliochama sp. Müllerried (1932b) Oax, Ejutla Biradiolites lombricalis d’Orb. sp. ?Alencasterites mooretownensis (Trechmann, 1924) Müllerried (1933a) Chis, Ocozocoautla Plagioptychus Arnaudi H. Douvillé, 1888 Plagioptychus muellerriedi Alencáster (1971) Chis, Vega del Paso Plagioptychus Toucasi Math., 1842 Plagioptychus antillarum (Douvillé, 1927) Müllerried (1933b) Chis Bournonia aff. africana H. Douvillé 1911 ? Radiolites cf. galloprovincialis Math. 1842 ? Radiolites cf. subradiosus Toucas 1908 ? (continued on next page) 10 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Sauvagesia cf. Da Rio Catullo, 1834 ? Distefanella lumbricalis Parona, 1900 ?Alencasterites mooretownensis (Trechmann, 1924) Plagioptychus Arnaudi nov. var. chiapasensis ?Plagioptychus antillarum (Douvillé, 1927) Radiolites lusitanicus Bayle, 1857 ? Hippurites (Vaccinites Fischer) giganteus ? d’Hombres-Firmas var. major Toucas, 1904 SLP Hippurites (Hippuritella H. Douvillé) cf. incisus ?Caribbea muellerriedi (Vermunt, 1937) (H. Douvillé) Toucas 1908 Oax Biradiolites lumbricalis ?Alencasterites mooretownensis (Trechmann, 1924) Müllerried (1934) Chis, [?Ocozocoautla] Sauvagesia (o Durania) n. sp. ?Thyrastylon? nicholasi (Whitfield, 1897a) Müllerried (1936a) Mex No details about Barrettia species Müllerried (1936b) Chis, Turonian Distefanella lombricalis (d’Orb.) Douvillé ?Alencasterites mooretownensis (Trechmann, 1924) Sauvagesia cf. da Rio Catullo ? Radiolites aff. lusitanicus Bayle ? Mex, Apaxco, Turonian Apricardia sp. Apricardia sp. Hippurites resectus var mexicana Hippurites resectus Defrance (1821) SLP, Huasteca, San Felipe beds, Turonian Sauvagesia degolyeri Stanton ?Radiolites degolyeri (Stanton, 1921) Chis, Turonian Barrettia ?Barrettia sp. Plagioptychus Toucasi Math. ?Plagioptychus antillarum (Douvillé, 1927) Chis, Coniacian Chiapasella radiolitiformis (Trechmann) Chiapasella radiolitiformis (Trechmann, 1924) Müllerried Pseudobarrettia ?Praebarrettia sparcilirata (Whitfield, 1897b) Biradiolites canaliculatus d’Orb. ? Durania cf. austinensis ? Chis, Coniacian-Santonian Radiolites cf. subradiosus Toucas ? Chis, Santonian Vaccinites giganteus d’Hombres-Firmas var. ? major Toucas Radiolites cf. galloprovincialis Mathéron ? Chis, Campanian Praeradiolites aff. subcoquandi Toucas ? Above Coralliochama ?Coralliochama sp. Biradiolites Aguilerae Boese Biradiolites aguilerae Böse (1906) Biradiolites cardenasensis Boese Biradiolites cardenasensis Böse (1906) Jones (1938) UMMP Coah, Sierra de Santa Ana, Sierra de la Peña Sauvagesia kellyi n. sp. ?Durania arnaudi Choffat (1886-1902) 16839, 16840–47, 16901 Müllerried (1942) W Chis, Campanian Coralliochama ?Coralliochama sp. Biradiolites aguilerae Böse Biradiolites aguilerae Böse (1906) Biradiolites cardenasensis Böse Biradiolites cardenasensis Böse (1906) Praeradiolites aff. subcoquandi Toucas ? W Chis, Santonian Vaccinites giganteus d’Hombres-Firmas var. ? major Toucas Radiolites cf. galloprovincialis Mathéron ? Radiolites cf. subradiosus Toucas ? W Chis, Coniacian Chiapasella radiolitiformis (Trechmann) Chiapasella radiolitiformis (Trechmann, 1924) Müllerried Pseudobarrettia ?Praebarrettia sparcilirata (Whitfield, 1897b) Biradiolites canaliculatus d’Orb. ? Durania cf. austinensis (Roemer) ? Radiolites cf. subradiosus Toucas ? W Chis, upper Turonian Distefanella lombricalis (d’Orb.) Douvillé ?Alencasterites mooretownensis (Trechmann, 1924) Sauvagesia cf. da-rio Toucas ? Radiolites aff. lusitanicus Bayle ? Apricardia sp. Apricardia sp. Sauvagesia degolyeri Stanton ?Radiolites degolyeri (Stanton, 1921) Barrettia Barrettia sp. Plagioptychus toucasi Math. ?Plagioptychus antillarum (Douvillé, 1927) Müllerried (1946) Ver, Córdoba, Cerro de Peñuela Neoradiolites ordoñezi n. sp. ? (undeterminable sections on building slabs) Müllerried (1947a) BC Coralliochama orcutti White Coralliochama orcutti White (1885) SLP, Cárdenas Coralliochama G. boehmi Boese Coralliochama gboehmi Böse (1906) Tamps, Rayón [ = San Antonio Rayón] Coralliochama n. sp. ?Coralliochama sp. Chis, Vega del Paso Plagioptychus toucasianus Plagioptychus antillarum (Douvillé, 1927) Chis, Ocozocoautla Plagioptychus arnaudi Plagioptychus muellerriedi Alencáster (1971) Müllerried (1947b) Ver, Córdoba, Cerro Peñuela Apricardia mongesi n. sp. ? (undeterminable sections on building slabs) Hippurites (Vaccinites) boehmi Douvillé pars. ?Vaccinites macgillavryi Palmer (1933) Bournonia carrilloi n. sp. ? (undeterminable sections on building slabs) Durania mexicana n. sp. ? (undeterminable sections on building slabs) Neoradiolites ordoñezi n. sp. ? (undeterminable sections on building slabs) Müllerried (1948a) Hgo, Valle del Mezquital, W of Tepé Hippurites llamasi n. sp. ? (undeterminable radiolitid) (continued on next page) 11 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Müllerried (1948b) Tab, Pasomono Durania austinensis (Roemer) Parona ? (undeterminable radiolitid fragment of right valve) Müllerried (1950a) NW Oax, SW Nochislán [ = Nochixtlán] Titanosarcolites Titanosarcolites sp. Barrettia monilifera Woodward, 1862 Barrettia monilifera Woodward (1862) Oax, W of Ejutla Distefanella lombricalis (d’Orbigny) 1847, ?Alencasterites mooretownensis (Trechmann, 1924) Douvillé, 1913 Gro, SE of Chilpancingo Distefanella lombricalis (d’Orbigny) 1847, ?Alencasterites mooretownensis (Trechmann, 1924) Douvillé, 1913 NL, Cerro de la Silla, Monterrey Chiapasella pauciplicata Müllerried, 1931 ?Chiapasella pauciplicata Müllerried (1931b) Müllerried (1950b) Chih, El Sotolar, Manuel Benavides Durania gonzalezi n. sp. ?Durania arnaudi Choffat (1886-1902) Müllerried (1950c) Mex, Apaxco [+Hgo] Hippurites (Hippuritella) resectus var. Hippurites resectus Defrance (1821) mexicana (Bárcena, 1875) Müllerried, 1930 Hippurites (Hippuritella) calamitiformis Hippurites resectus Defrance (1821) Bárcena, 1875 Müllerried (1950d) Mor, Tlaquiltenango, Los Hornos Durania cornu-pastoris (Des Moulins) Parona ?Durania arnaudi Choffat (1886-1902) pars. Müllerried (1952) Coah Durania coahuilensis n. sp. ?Durania arnaudi Choffat (1886-1902) Müllerried (1953) Coah, Hendidura [ = Indidura] Formation Durania kellyi (Jones) pars ?Durania arnaudi Choffat (1886-1902) Durania cf. cornupastoris (Desmoulins) ?Durania arnaudi Choffat (1886-1902) Parona Durania cf. D. gonzalezi ?Durania arnaudi Choffat (1886-1902) Durania kellyi (Jones) sp. ?Durania arnaudi Choffat (1886-1902) Hippurites (Hippuritella H. Douvillé; Orbignya Hippurites resectus Defrance (1821) Toucas) resectus var. mexicana (Bárcena) Müllerried, 1930 Bauman (1958) NMNH Mor, Cuautla Formation Radiolites newelli n. sp. ?Eoradiolites liratus (Conrad, 1852) 27198 Radiolites muellerriedi n. sp. ?Eoradiolites liratus (Conrad, 1852) 27199 Chubb (1959) Central Chis, Carretera Formation Biradiolites ssp. ?Biradiolites sp. Caprinidae ?Caprinidae indet. Central Chis, Campeche Beds Chiapasella radiolitiformis Chiapasella radiolitiformis (Trechmann, 1924) Titanosarcolites ?Titanosarcolites sp. Durania nicholasi Thyrastylon? nicholasi (Whitfield, 1897a) Central Chis, Nuevo Beds Plagioptychus aff. P. toucasi Matheron ?Plagioptychus antillarum (Douvillé, 1927) Coralliochama aff. C. g. boehmi Böse ?Plagioptychus muellerriedi Alencáster (1971) Central Chis, San Luis Conglomerate “Pseudobarrettia” ?Praebarrettia sparcilirata (Whitfield, 1897b) Central Chis, Piedra Parada Beds ? Barrettia ?Barrettia sp. Central Chis, Sierra Madre Limestone Distefanella lombricalis ?Alencasterites mooretownensis (Trechmann, 1924) (uppermost part) Sauvagesia ssp. ?Sauvagesia sp. Durania ssp. ?Durania sp. Damestoy (1965) MNHN SC, 19 km S of Ensenada Coralliochama flouriei n. sp. Coralliochama orcutti White (1885) Myers (1968) TMM SLP, Cardenas Formation Coralliochama gboehmi Böse Coralliochama gboehmi Böse (1906) 15567, 15582, 15583 Hippurites muellerriedi (Vermunt) Caribbea muellerriedi (Vermunt, 1937) 15568, 15584 Hippurites perkinsi n. sp. Caribbea muellerriedi (Vermunt, 1937) 15569 Biradiolites aguilerae Böse Biradiolites aguilerae Böse, 1906 15560, 15580 Biradiolites cardenasensis Böse Biradiolites cardenasensis Böse, 1906 15511–15513 Tampsia floriformis n. sp Tampsia floriformis Myers (1968) 14925, 14955 Tampsia poculiformis n. sp. Tampsia floriformis Myers (1968) 14921, 14933 Durania ojanchalensis Myers, 1968 Huasteca ojanchalensis (Myers, 1968) [? = Huasteca 14996, 14984 huasteca (Adkins, 1930)] Alencáster (1971) IGM Chis, Vega del Paso Plagioptychus cf. P. jamaicensis (Whitfield) Plagioptychus antillarum (Douvillé, 1927) 2130–2133 Chis, Ocozocoautla Plagioptychus muellerriedi n. sp. Plagioptychus muellerriedi Alencáster (1971) 2134–2141 Chis, Ocozocoautla [?] Plagioptychus agariciformis n. sp. Coralliochama gboehmi Böse (1906) [probably from 2124,2125 Cárdenas] Chis, Ocuilapa Titanosarcolites giganteus (Whitfield) pars Titanosarcolites giganteus (Whitfield, 1897a); pars 2173–2178 Alencasteria macrotubularis Mitchell (2013); pars Caenosarcolites oddsensis (Stephenson, 1938) Chis, Vega del Paso Titanosarcolites macgillavryi n. sp. Stellacaprina macgillavryi (Alencáster, 1971) 2173–2178 Chis, Ocozocoautla Biradiolites rudissimus Trechmann Trechmannites rudissimus (Trechmann, 1924) 2092–2097 Chis, Vega del Paso Biradiolites cancellatus (Whitfield) Bournonia baileyi Chubb (1967) 2105–2109 Chis, Vega del Paso Bournonia cardenasensis (Böse) Biradiolites cardenasensis Böse (1906) 2110–2114 Chis, Berriozábal Thyrastylon chubbi n. sp. Thyrastylon adhaerens (Whitfield, 1897a) 2088–2091 Chis, Ocozocoautla Sauvagesia macroplicata (Whitfield) “Radiolites” macroplicata Whitfield (1897a) 2084–2087 Chis, Ocozocoautla Durania ojanchalensis Myers 2127–2129 (continued on next page) 12 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Chis, Ocozocoautla [?] Tampsia floriformis Myers Tampsia floriformis Myers (1968) [probably from 2102–2104 Cárdenas] Chis, Ocozocoautla Durania nicholasi (Whitfield) Thyrastylon? nicholasi Whitfield (1897a) 2098–2101 Chis, Vega del Paso Barrettia monilifera Woodward Barrettia monilifera Woodward (1862) 2150–2156 Chis, Vega del Paso Barrettia multilirata (Whitfield) pars Barrettia monilifera Woodward (1862); pars Barrettia 2142–2146 gigas Chubb (1955); pars Parastroma trechmanni Chubb (1967) Chis, Vega del Paso Barrettia gigas Chubb pars Barrettia gigas Chubb (1955); pars Parastroma 2147–2149 trechmanni Chubb (1967) Chis, Ocozocoautla Praebarrettia sparcilirata (Whitfield) Praebarrettia sparcilirata (Whitfield, 1897b) 2158–2166 Chis, Ocozocoautla Chiapasella radiolitiformis (Trechmann) Chiapasella radiolitiformis (Trechmann, 1924) 2115-2123, 2185 Marincovich (1975) LACM BC, Punta Banda, Rosario Formation Coralliochama orcutti White, 1885 Coralliochama orcutti White (1885) Young (1984) SLP, W Santa Catarina, Cuautla Formation Hippurites resectus Defrance, 1821 var. Hippurites resectus Defrance (1821) mexicana Barcena, 1875 SLP, San Francisco, Valle de Fantasmas, Radiolites sp. ? Cuautla Formation SLP, Charco Blanco, Cuautla Formation Sauvagesia sp. ? Durania sp. ? Jal, Cementos Tolteca quarry Radiolites sp. cf. R. newelli Bauman, 1958 ?Eoradiolites liratus (Conrad, 1852) Alencáster et al. (1987) IGM Gro, La Esperanza, Almolonga-Hueytalpan Vaccinites praegiganteus Toucas, 1904 Vaccinites macgillavryi Palmer (1933) 4309–4310 Vaccinites grossouvrei Douvillé, 1894 Vaccinites macgillavryi Palmer (1933) 4311 Vaccinites gosaviensis Douvillé, 1890 Vaccinites macgillavryi Palmer (1933) 4312 Vaccinites giganteus d’Hombre-Firmas (1838) Vaccinites macgillavryi Palmer (1933) 4313–4314 Vaccinites sp. Vaccinites macgillavryi Palmer (1933) 4315 Waite (1987) Chis, Sierra Madre Limestone Sauvagesia sp. ?Sauvagesia sp. Perrilliat (1989) Mexico all Mexican types up to date Alencáster and Michaud (1990) IGM Chis, Suchiapa Formation Vaccinites inaequicostatus vermunti Mac Vaccinites vermunti MacGillavry, 1937 4564–4566 Gillavry Durania curasavica (Martin) ?Potosites tristantorresi Alencáster and Pons, in Pons et al., 4567 2010 Aranda-Manteca (1991) BC, Punta Banda, Rosario Formation Coralliochama orcutti White, 1885 Coralliochama orcutti White (1885) Alencáster and Pons (1992) IGM Chis, Angostura Formation “Radiolites” macroplicatus Whitfield, 1897 “Radiolites” macroplicatus Whitfield (1897a) 2087bis, 6193, 6194 Chis, Ocozocoautla Formation Chiapasella radiolitiformis (Trechmann, Chiapasella radiolitiformis (Trechmann, 1924) 2118, 2184, 6196- 1924) 6198 García-Barrera et al. (1998) IGM/FCMP Chis, Angostura Formation Antillocaprina trilobata García Barrera and pars Rotacaprina? trilobata García Barrera and Avendaño, Avendaño, 1998 in García Barrera et al., 1998; pars Antillocaprina suboccidentalis Chubb (1967) Coralliochama gboehmi Böse Coralliochama gboehmi Böse (1906) [?from Cárdenas] 2124,2125 Alencáster et al. (1999a) IGM SLP, Cardenas Formation, Amoladeras Praebarrettia sparcilirata (Whitfield) Praebarrettia armini (Götz and Mitchell, 2009) Coralliochama gbohemi Böse Coralliochama gbohemi Böse, 1906 Bournonia cardenasensis (Böse) Biradiolites cardenasensis Böse (1906) Durania ojanchalensis Myers Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Biradiolites rudissimus Trechmann Trechmannites rudissimus (Trechmann, 1924) SLP, Cardenas Formation, Cárdenas Durania ojanchalensis Myers Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Bournonia cardenasensis (Böse) Biradiolites cardenasensis Böse (1906) Radiolites macroplicatus Whitfield Coralliochama gbohemi Böse (1906) Coralliochama gbohemi Böse Coralliochama gbohemi Böse (1906) Tampsia floriformis Myers Tampsia floriformis Myers (1968) Alencáster et al. (1999b) IGM SLP, San Luis-Matehuala Hippurites resectus mexicana Bárcena Hippurites resectus Defrance (1821) Eoradiolites cf. liratus (Conrad) Eoradiolites liratus Conrad (1852) Durania austinensis (Römer) Potosites tristantorresi Alencáster and Pons, in Pons et al., 2010 Vaccinites sp. Vaccinites macgillavryi Palmer (1933) Alencáster et al. (2001) SLP,Cárdenas; Chiapas Tampsia Tampsia sp. Praebarrettia Praebarrettia sp. Radiolites Radiolites sp. Bournonia Bournonia sp. Durania Durania sp. Thyrastylon Thyrastylon sp. Coralliochama Coralliochama sp. (continued on next page) 13 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Hippurites Hippurites sp. Biradiolites Biradiolites sp. Plagioptychus Plagioptychus sp. Sauvagesia Sauvagesia sp. Alencáster and Omaña (2005) SLP, Cárdenas Formation Biradiolites cardenasensis Böse Biradiolites cardenasensis Böse (1906) Biradiolites rudissimus Trechmann Trechmannites rudissimus (Trechmann, 1924) Radiolites macroplicata Whitfield ?Coralliochama gbohemi Böse (1906) Durania ojanchalensis Myers Huasteca ojanchalensis (Myers, 1968) [?Huasteca huasteca (Adkins, 1930)] Tampsia floriformis Myers Tampsia floriformis Myers (1968) Tampsia bishopi Stephenson ?Tampsia bishopi Stephenson (1922) Thyrastylon adherens (Whitfield) ? Coralliochama gbohemi Böse Coralliochama gbohemi Böse (1906) Praebarrettia sparcilirata (Whitfield) Praebarrettia armini (Götz and Mitchell, 2009) Chis, Angostura Formation Titanosarcolites Titanosarcolites sp. Barrettia Barrettia sp. Macgillavrya ?Thyrastylon? sp. Parastroma Parastroma sp. Antillocaprina Antillocaprina sp. Plagioptychus Plagioptychus sp. Filkorn et al. (2005) Chis, Angostura Formation Sauvagesia macroplicata (Whitfield, 1897) “Radiolites” macroplicata Whitfield (1897a) Götz et al. (2005) N BC, Rosario Formation Coralliochama orcutti White Coralliochama orcutti White (1885) Oviedo (2006) IGM/PUAB SLP, Cárdenas, Amoladeras, Cardenas Biradiolites rudissimus Trechmann, 1924 Trechmannites rudissimus (Trechmann, 1924) Formation SLP, Cárdenas, Amoladeras, Cardenas Bournonia cardenasensis Böse, 1906 Biradiolites cardenasensis Böse, 1906 Formation SLP, Llano las hormigas, (El Abra Formation) Durania arnaudi (Choffat, 1891) Durania arnaudi Choffat (1886-1902) Gro, Almolonga-Hueyitlalpan, Cuautla Durania curasavica (Martin, 1885) Durania curasavica Martin (1885) Formation SLP, Amoladeras, Cardenas Formation Durania ojanchalensis Myers, 1968 Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Gro, Temalac, Cuautla Formation Durania nov. sp. Durania sp. SLP, Cerritos, (Cardenas Formation) Durania? sp. Durania? sp. SLP, La Calera, El Doctor Formation; Durania sp. Durania sp. SLP, Llano las hormigas, (El Abra Formation) Eoradiolites liratus (Conrad, 1852) Eoradiolites liratus (Conrad, 1852) SLP, Temazcal; Cantera Cerro El Cristito; El Radiolites acutocostata (Adkins, 1930) ?Radiolites degolyeri (Stanton, 1921) Aguaje, Guadalcázar, (Cardenas Formation) SLP, Cárdenas, Cardenas Formation Tampsia floriformis Myers, 1968 Tampsia floriformis Myers (1968) Chis, Malpaso road (Ocozocoautla Formation) Thyrastylon adhaerens (Whitfield, 1897) Thyrastylon adhaerens (Whitfield, 1897a) Chis, Ocozocoautla; Río Suchiapa, “Thyrastylon” nicholasi (Whitfield, 1897) Thyrastylon? nicholasi (Whitfield, 1897a) Ocozocoautla Formation SLP, Temaxcal; Cantera Cerro El Cristito, Radiolitidae nov. gen. Potosites tristantorresi Alencáster and Pons, in Pons et al., (Cardenas Formation) 2010 SLP, Cárdenas, Cardenas Formation Coralliochama gboehmi Böse, 1906 Coralliochama gboehmi Böse (1906) SLP, Cárdenas; Amoladeras, Cardenas Mitrocaprina tschoppi (Palmer, 1933) Mitrocaprina tschoppi (Palmer, 1933) Formation Chis, Ocozocoautla, Ocozocoautla Formation Plagioptychus muellerriedi Alencáster, 1971 Plagioptychus muellerriedi Alencáster (1971) SLP, La Calera, El Doctor Formation; Llano las Hippurites resectus Defrance, 1821 Hippurites resectus Defrance (1821) hormigas, (El Abra Formation) SLP, Amoladeras; Cardenas Formation; Praebarrettia sparcilirata (Whitfield, 1897) pars Praebarrettia armini Götz and Mitchell (2009); pars Chiapas, Malpaso road (Angostura Praebarrettia sparcilirata Whitfield (1897b) Formation) SLP, Sierra de la Ruda, (El Abra Formation); Vaccinites macgillavryi Palmer, 1933 Vaccinites macgillavryi Palmer (1933) Gro., La Esperanza; Almolonga- Hueyitlalpan, Cuautla Formation Chis, Cantera rancho El Comiteco, (Angostura Vaccinites vermunti Mac Gillavry, 1937 Vaccinites vermunti MacGillavry, 1937 Formation) SLP, Temaxcal; Cantera Cerro El Cristito, Vaccinites nov. sp. Vaccinites vermunti MacGillavry, 1937 (Cardenas Formation) Chis, Malpaso road (Ocozocoautla Formation) Antillocaprina pugniformis (Palmer, 1933) Antillocaprina quadrangularis (Whitfield, 1897a) Chis, Malpaso road (Ocozocoautla Formation) Titanosarcolites giganteus (Whitfield, 1897) pars Titanosarcolites giganteus (Whitfield, 1897a); pars Alencasteria macrotubularis Mitchell (2013) Chis, Vega del Paso Titanosarcolites macgillavryi Alencáster, 1971 Stellacaprina macgillavryi (Alencáster, 1971) Chis, Malpaso road, rancho La Peregrina Titanosarcolites nov. sp. Alencasteria macrotubularis Mitchell (2013) (Ocozocoautla Formation) Schafhauser and Karlsruhe (2006) SLP, Cardenas Formation Mitrocaprina cf. tschoppi Palmer, 1933 ?Mitrocaprina tschoppi Palmer (1933) Coralliochama gboehmi Böse, 1906 Coralliochama gboehmi Böse (1906) Macgillavryia nicholasi (Whitfield, 1897) ?Thyrastylon? nicholasi Whitfield (1897a) Bournonia cardenasensis (Böse, 1906) Biradiolites cardenasensis Böse (1906) Tampsia poculiformis Myers, 1968 Tampsia floriformis Myers (1968) Hippurites perkinsi Myers, 1968 Caribbea muellerriedi (Vermunt, 1937) (continued on next page) 14 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Praebarrettia? sparcilirata (Whitfield, 1897) Praebarrettia armini (Götz and Mitchell, 2009) Oviedo et al. (2007) IGM/PUAB SLP Hippurites resectus Defrance, 1821 Hippurites resectus Defrance (1821) Durania arnaudi (Choffat, 1891) Durania arnaudi Choffat (1886-1902) SLP; Gro. Vaccinites macgillavryi Palmer, 1933 Vaccinites macgillavryi Palmer (1933) Durania curasavica (Martin, 1885) Durania curasavica Martin (1885) SLP, Non-designed massive limestones unit Vaccinites n. sp. Vaccinites vermunti MacGillavry, 1937 Barrettia monilifera Woodward, 1862 Barrettia ruseae Chubb (1967) Torreites sanchezi (Douvillé, 1927) Torreites sanchezi (Douvillé, 1927) Radiolites acutocostata (Adkins, 1930) ?Radiolites degolyeri (Stanton, 1921) SLP, Cardenas Formation (lower) Praebarrettia sparcilirata (Whitfield, 1897) Praebarrettia armini (Götz and Mitchell, 2009) Biradiolites rudissimus Trechmann, 1924 Trechmannites rudissimus (Trechmann, 1924) Bournonia cardenasensis (Böse, 1906) Biradiolites cardenasensis Böse (1906) Durania ojanchalensis Myers, 1968 Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Mitrocaprina tschoppi (Palmer, 1933) Mitrocaprina tschoppi (Palmer, 1933) SLP, Cardenas Formation (upper) Hippurites muellerriedi Vermunt, 1937 Caribbea muellerriedi (Vermunt, 1937) Tampsia floriformis Myers, 1968 Tampsia floriformis Myers (1968) Schafhauser and GötzStinnesbeck (2007) SLP, Cardenas Formation Mitrocaprina cf. tschoppi Palmer ?Mitrocaprina tschoppi Palmer (1933) Coralliochama gboehmi Böse Coralliochama gboehmi Böse (1906) Durania ojanchalensis Myers Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Macgillavryia nicholasi (Whitfield) ?Thyrastylon? nicholasi Whitfield (1897a) Biradiolites aguilerae Böse Biradiolites aguilerae Böse (1906) Bournonia cardenasensis Böse Biradiolites cardenasensis Böse (1906) Tampsia poculiformis Myers Tampsia floriformis Myers (1968) Tampsia floriformis Myers Tampsia floriformis Myers (1968) Hippurites perkinsi Myers Caribbea muellerriedi (Vermunt, 1937) Hippurites muellerriedi (Vermunt) Caribbea muellerriedi (Vermunt, 1937) ?Praebarrettia sparcilirata (Whitfield) Praebarrettia armini (Götz and Mitchell, 2009) Pichardo (2008) PUAB SLP, Non-designed limestones unit Biradiolites sp. Biradiolites sp. Durania sp. Durania sp. Radiolites acutocostata (Adkins, 1930) ?Radiolites degolyeri (Stanton, 1921) Radiolitidae nov. gen. Potosites tristantorresi Alencáster and Pons, in Pons et al., 2010 Mitrocaprina sp. Mitrocaprina sp. Plagioptychus sp. Plagioptychus sp. Barrettia monilifera Woodward, 1862 Barrettia ruseae Chubb (1967) Torreites sanchezi (Douvillé, 1927) Torreites sanchezi (Douvillé, 1927) Vaccinites sp. Vaccinites vermunti MacGillavry, 1937 Götz and Mitchell (2009) IGM SLP, Cardenas Formation Laluzia armini Götz and Mitchell, 2009 Praebarrettia armini (Götz and Mitchell, 2009) 9023,9017, 9020–9022, 8888 Pons et al. (2010) IGM/PUAB/FCMP SLP, Temaxcal limestones Potosites tristantorresi Alencáster and Pons Potosites tristantorresi Alencáster and Pons, in Pons et al. 09/032 (2010) Radiolites acutocostata (Adkins, 1930) ?Radiolites degolyeri (Stanton, 1921) Barrettia cf. ruseae Chubb, 1967 Barrettia cf. ruseae Chubb (1967) 1288 719 Torreites sanchezi (Douvillé, 1927) Torreites sanchezi (Douvillé, 1927) 1286 712, 81000 Vaccinites vermunti Mac Gillavry, 1937 Vaccinites vermunti MacGillavry, 1937 0919 684 = 9537 Plagioptychus sp. Plagioptychus sp. 1286 706 Plagioptychidae genus indeterminate A Plagioptychidae genus indeterminate 81016 Plagioptychidae genus indeterminate B Plagioptychidae genus indeterminate 9538 Antillocaprinidae genus indeterminate Antillocaprinidae genus indeterminate Mitchell and Pons (2010) TMM Tamps, Mendez Formation Chiapasella aguilae (Adkins, 1930) Chiapasella aguilae (Adkins, 1930) BEG2066.5 Mitchell (2013) IGM Chis Caenosarcolites oddsensis (Stephenson, 1938) Caenosarcolites oddsensis (Stephenson, 1938) Antillocaprina suboccidentalis Chubb, 1968 Antillocaprina suboccidentalis Chubb (1967) Rotacaprina? trilobata (García-Barrera and Rotacaprina? trilobata (García-Barrera and Avendaño, in Avendaño) García Barrera et al., 1998) Alencasteria macrotubularis Mitchell, 2013 Alencasteria macrotubularis Mitchell (2013) Stellacaprina macgilavryi (Alencáster, 1971) Stellacaprina macgilavryi (Alencáster, 1971) ? Oligosarcolites monotubularis Mitchell and ? Oligosarcolites monotubularis Mitchell and Gunter (2006) Gunter, 2006 Pons et al. (2013) IGM/PUAB SLP, Cardenas Formation Biradiolites aguilerae Böse, 1906 Biradiolites aguilerae Böse (1906) Biradiolites cardenasensis Böse, 1906 Biradiolites cardenasensis Böse (1906) Huasteca ojanchalensis (Myers, 1968) Huasteca ojanchalensis (Myers, 1968) [? = Huasteca huasteca (Adkins, 1930)] Tampsia floriformis Myers, 1968 Tampsia floriformis Myers (1968) Trechmannites rudissimus (Trechmann, 1924) Trechmannites rudissimus (Trechmann, 1924) Caribbea muellerriedi (Vermunt, 1937) Caribbea muellerriedi (Vermunt, 1937) Praebarrettia armini (Götz and Mitchell, 2009) (continued on next page) 15 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 (continued ) Author/Localities Reported taxa Current identification Repository Praebarrettia sparcilirata (Whitfield, 1897) sensu lato Coralliochama gboehmi Böse, 1906 Coralliochama gboehmi Böse (1906) Mitrocaprina tschoppi (Palmer, 1933) Mitrocaprina tschoppi (Palmer, 1933) Pons et al. (2016) IGM/PUAB/FCMP Chis, Suchiapa Formation (Angostura Block) Alencasteria macrotubularis Mitchell Alencasteria macrotubularis Mitchell (2013) Stellacaprina? macgillavryi (Alencáster) Stellacaprina? macgillavryi (Alencáster, 1971) Antillocaprinidae indet. Antillocaprinidae indet. “Postcaprinula” Alencáster (in collection) “Postcaprinula” Alencáster (in collection IGM) Barrettia monilifera Woodward, Barrettia monilifera Woodward (1862) Whitfieldiella gigas (Chubb) Barrettia gigas (Chubb, 1955) Parastroma trechmanni Chubb Parastroma trechmanni Chubb (1967) Plagioptychus sp. Plagioptychus antillarum (Douvillé, 1927) Mitrocaprina sp. Mitrocaprina sp. Alencasterites mooretownensis (Trechmann) Alencasterites mooretownensis (Trechmann, 1924) Bournonia baileyi Chubb, Bournonia baileyi Chubb (1967) Biradiolites? sp. Biradiolites? sp. Thyrastylon? nicholasi (Whitfield) Thyrastylon? nicholasi Whitfield (1897a) Chis, Suchiapa Formation (Tuxtla Gutiérrez Antillocaprinidae indet. Antillocaprinidae indet. Block) Barrettia sp. cf. B. ruseae Chubb Barrettia sp. cf. B. ruseae Chubb (1967) Vaccinites vermunti Mac Gillavry Vaccinites vermunti MacGillavry, 1937 Radiolites acutocostata (Adkins) ?Radiolites degolyeri (Stanton, 1921) Potosites tristantorresi Alencáster and Pons Potosites tristantorresi Alencáster and Pons, in Pons et al., 2010 Chiapasella sp. Chiapasella sp. Chis, Ocozocoautla Formation Praebarrettia sparcilirata (Whitfield) Praebarrettia sparcilirata (Whitfield, 1897b) Plagioptychus muellerriedi Alencáster Plagioptychus muellerriedi Alencáster (1971) Biradiolites cardenasensis Böse Biradiolites cardenasensis Böse (1906) Chiapasella radiolitiformis (Trechmann) Chiapasella radiolitiformis (Trechmann, 1924) Huasteca ojanchalensis (Myers) Huasteca ojanchalensis (Myers, 1968) [?=Huasteca huasteca (Adkins, 1930)] Thyrastylon? nicholasi (Whitfield) Thyrastylon? nicholasi Whitfield (1897a) Trechmannites rudissimus (Trechmann) Trechmannites rudissimus (Trechmann, 1924) Chis, Angostura Formation Antillocaprina pugniformis (Palmer) Antillocaprina quadrangularis Whitfield (1897a) Antillocaprina trilobata (García Barrera and pars Rotacaprina? trilobata García Barrera and Avendaño, Avendaño) in García Barrera et al., 1998; pars Antillocaprina suboccidentalis Chubb (1967) Caenosarcolites oddsensis (Stephenson) Caenosarcolites oddsensis (Stephenson, 1938) Caenosarcolites scholaris Mitchell ? Eosarcolites radiatus Mitchell ? Titanosarcolites giganteus (Whitfield) Titanosarcolites giganteus Whitfield (1897a) Antillocaprinidae indet. Antillocaprinidae indet. Plagioptychus sp. Plagioptychus fragilis Chubb (1967) Mitrocaprina sp. Mitrocaprina tschopi (Palmer, 1933) “Radiolites” macroplicatus Whitfield “Radiolites” macroplicatus Whitfield (1897a) Biradiolites cardenasensis Böse Biradiolites cardenasensis Böse (1906) Biradiolites? sp. Biradiolites? sp. Bournonia cancellata (Whitfield) Bournonia cancellata Whitfield (1897a) Chiapasella radiolitiformis (Trechmann) Chiapasella radiolitiformis (Trechmann, 1924) Thyrastylon adhaerens (Whitfield) Thyrastylon adhaerens Whitfield (1897a) Pons et al. (2017) IGM/PUAB/FCMP Chis, Suchiapa Formation (Angostura Block) Plagioptychus antillarum (Douvillé, 1927) Plagioptychus antillarum (Douvillé, 1927) Mitrocaprina sp. Mitrocaprina sp. Chis, Ocozocoautla Formation Plagioptychus muellerriedi Alencáster, 1971 Plagioptychus muellerriedi Alencáster (1971) Chis, Angostura Formation (Tuxtla Gutiérrez Plagioptychus fragilis Chubb, 1967 Plagioptychus fragilis Chubb (1967) Block) Mitrocaprina tschoppi (Palmer, 1933) Mitrocaprina tschoppi (Palmer, 1933) Pons et al. (2019) IGM/PUAB/FCMP Chis, Suchiapa Formation (Tuxtla Gutiérrez Vaccinites vermunti Mac Gillavry, 1937 Vaccinites vermunti MacGillavry, 1937 Block) Barrettia cf. ruseae Chubb, 1955 Barrettia cf. ruseae Chubb (1967) Chis, Suchiapa Formation (Angostura Block) Barrettia monilifera Woodward, 1862 Barrettia monilifera Woodward (1862) Barrettia gigas Chubb, 1955 Barrettia gigas Chubb (1955) Parastroma trechmanni Chubb, 1967 Parastroma trechmanni Chubb (1967) Chis, Ocozocoautla Formation Praebarrettia sparcilirata (Whitfield, 1897) Praebarrettia sparcilirata (Whitfield, 1897b) Mexican States abbreviations: BC=Baja California; Chih = Chihuahua; Chis = Chiapas; Coah = Coahuila; Gro = Guerrero; Jal = Jalisco; Mex = México; Mich = Michoacan; Mor = Morelos; NL=Nuevo León; Qro = Querétaro; SLP=San Luis Potosí; Tamps = Tamaulipas; Ver = Veracruz Institutional repositories abbreviations in text: 2. Material and methods. Not visited collections are indicated in bold. References * Adkins, W.S., 1930. New rudistids from the Texas and Mexican cretaceous. The University of Texas Bulletin, Contributions to Geology 3001 (77–100), 121–133. Aguayo Camargo, J.E., 1978. Sedimentary environments and diagénesis of a Cretaceous * Adkins, W.S., 1928. Handbook of Texas Cretaceous Fossils. The University of Texas reef complex, eastern Mexico. Anales Centro Ciencias del Mar y Limnología UNAM 5 Bulletin. (1), 83–140. 16 J.M. Pons et al. Journal of South American Earth Sciences 109 (2021) 103237 * Alencáster, G., 1971. Rudistas del Cretácico Superior de Chiapas. Parte I. Palaontol. d’Orbigny, A., 1847. Paléontologie française. Terrains crétacés, 4, 490–599 [Rudists Mexic. 34, 1–91. part, pp. 157–267, pl. 526-599, was published in 1850]. Alencáster, G., García-Barrera, P., 2008. Albian radiolitid rudists (Mollusca Bivalvia) Douvillé, H., 1888. Études sur les Caprines. Bull. Soc. Geol. Fr. 16 (3), 699–730. from east-Central Mexico. Geobios 41, 571–587. * Douvillé, H., 1891. Études sur les rudistes. Révision des principales espèces * Alencáster, G., Michaud, F., 1990. Rudistas (Bivalvia-Hippuritacea) del Cretácico d’Hippurites. Mémoires de la Société géologique de France, Paléontologie 1–6, superior de la región de Tuxtla Gutiérrez, Chiapas (México). Actas Facultad de 1–230. Mémoire No. 6. Ciencias de la Tierra UANL 4, 175–193. Douvillé, H., 1908. Sur la classification des Radiolitidés. Bull. Soc. Geol. Fr. 8, 308–310. * Alencáster, G., Omaña, L., 2005. Maastrichtian demise of rudists in the valles-san Luis Douvillé, H., 1909. Sur le genre Eoradiolites. Comptes Rendus des Séances de la Société Potosí and Chiapas platforms, Mexico. 17-18. In: Filkorn, H.F., Johnson, C.C., géologique de France (série 4) 9, 77. Molineux, A., Scott, R.W. (Eds.), Abstracts and Post-Congress Field Guide, Seventh Douvillé, H., 1910. Études sur les rudistes. Rudistes de Sicile, d’Algérie, d’Égypte, du International Congress on Rudists. SEPM Miscellaneous Publication Number 6. Liban et de la Perse. 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(Eds.), (Bivalvia-Hippuritacea) del Cretácico superior de la parte central del estado de Abstracts and Post-Congress Field Guide, Seventh International Congress on Rudists. Guerrero. Rev. Soc. Mexic. Palaontol. 1 (1), 24–39. SEPM Miscellaneous Publication Number 6. * Alencáster, G., Omaña, L., Pons, J.M., Torres, J.R., Tristán, M., Vázquez, N., 1999a. Fischer, P., 1887. In: Savy, F. (Ed.), Manuel de Conchyliologie et de Paléontologie Upper cretaceous rudists and larger foraminifera from the Cárdenas region, san Luis conchyliologique ou Histoire Naturelle des Mollusques vivants et fossiles, fasc, vol. Potosí, México. Erlanger Geol. Abh. 3, 6–7. 11, pp. 1009–1369 (Paris). * Alencáster, G., Torres-Hernández, J.R., Tristán-González, M., Barbosa-Gudiño, R., *García-Barrera, P., Avendaño Gil, J., Omaña, L., Alencáster, G., 1998. Antillocaprina López-Doncel, R., Pons, J.M., Omaña, L., 1999b. 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