c5987c0859c63a087232ca03834d8cd2 ICP02986.pdf 1a4011876220bab8edb11dfcb6004bd0111d7123 ICP02986.pdf 35de58d4e7051ec892dea099ca0abc2676d3273552d0b4c09fb52db9ffc5720d ICP02986.pdf Title: Growing giants: ontogeny and life history of the temnospondyl Mastodonsaurus giganteus (Stereospondyli) from the Middle Triassic of Germany Subject: The Middle Triassic capitosaur Mastodonsaurus giganteus was the largest temnospondyl and the dominating aquatic predator in many European freshwater to brackish ecosystems. It is represented by numerous size classes, which are described and analysed for the first time. The documented size range encompasses specimens between 12–15 mm and 1200 mm in skull length. Early growth stages are restricted to dentaries and interclavicles, whereas juveniles are represented by partial skulls, mandibles, and girdles. The smallest specimens already possessed diagnostic features of the taxon, and small juveniles also shared the dermal ornament with larger specimens. The heavy, disc-shaped intercentra were established early in the juvenile phase. Cranial proportions were remarkably conservative throughout ontogeny, with the orbits proportionately decreasing in size only very moderately, the postorbital skull becoming slightly longer and the occipital margin more concave in the largest forms. Analysis of frequency distributions of M. giganteus in different Lower Keuper deposits in southern Germany reflects habitat preferences in specific phases of its life cycle. The coal-bearing deposit at Gaildorf yielded unusually large specimens with relatively well-ossified appendicular skeletons. In the more common lake shore facies, only adult specimens are present. In turn, juveniles might have dwelled in calmer environments. Smaller lakes were apparently less attractive than larger or deeper water bodies that provided sufficient resources for several temnospondyls, and juvenile specimens have been identified from all of them. The diverse actinopterygian fish fauna provided prey for all growth stages of the large temnospondyl predators. Author: Rainer R. Schoch Creator: Adobe InDesign 20.0 (Windows) Producer: Adobe PDF Library 17.0 CreationDate: Mon Dec 23 15:59:53 2024 CET ModDate: Mon Dec 23 16:00:03 2024 CET Custom Metadata: no Metadata Stream: yes Tagged: yes UserProperties: no Suspects: no Form: none JavaScript: no Pages: 22 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 10615315 bytes Optimized: yes PDF version: 1.4 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- VRNHXX+TimesNewRomanPS-ItalicMT CID TrueType Identity-H yes yes yes 740 0 EXZETF+SymbolMT CID TrueType Identity-H yes yes yes 741 0 VJPMXX+TimesNewRomanPSMT CID TrueType Identity-H yes yes yes 742 0 BINRDL+NewsGothicStd Type 1C Custom yes yes yes 726 0 DYKYFH+FFMarkPro-Bold Type 1C WinAnsi yes yes yes 743 0 INBHXX+TimesNewRomanPSMT TrueType WinAnsi yes yes yes 727 0 MBCFBP+TimesNewRomanPS-ItalicMT TrueType WinAnsi yes yes yes 728 0 DQMDFH+NewsGothicMT TrueType WinAnsi yes yes yes 729 0 TEKQVB+TimesNewRomanPS-BoldMT TrueType WinAnsi yes yes yes 749 0 JPKILV+NewsGothicMT-Italic TrueType WinAnsi yes yes yes 750 0 BINRDL+NewsGothicStd Type 1C WinAnsi yes yes yes 69 0 JPKILV+NewsGothicMT-Italic TrueType WinAnsi yes yes yes 70 0 GNWQVB+TimesNewRomanPS-BoldItalicMT TrueType WinAnsi yes yes yes 73 0 BINRDL+NewsGothicStd Type 1C WinAnsi yes yes yes 74 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2025-02-17 16:57:24 CET RepresentationInformation: ICP02986.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2025-02-13 11:03:14 CET Size: 10615315 Format: PDF Version: 1.4 Status: Well-Formed, but not valid SignatureMatches: PDF-hul ErrorMessage: Annotation object is not a dictionary ID: PDF-HUL-114 Offset: 5759 MIMEtype: application/pdf Profile: Linearized PDF PDFMetadata: Objects: 791 FreeObjects: 586 IncrementalUpdates: 1 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Language: en-US Outlines: Item: Title: Growing giants: ontogeny and life history of the temnospondyl Mastodonsaurus giganteus (Stereospondyli) from the Middle Triassic of Germany Item: Title: Abstract Item: Title: Introduction Item: Title: Materials and methods Children: Item: Title: Material Item: Title: Ontogeny Item: Title: Abbreviations Item: Title: Systematic Palaeontology Children: Item: Title: Genus Mastodonsaurus Jaeger, 1828 Item: Title: Mastodonsaurus giganteus Jaeger, 1828 Item: Title: Ontogeny of Mastodonsaurus giganteus Children: Item: Title: Smallest growth stages Item: Title: Juveniles Item: Title: Adults Item: Title: Giant specimens Item: Title: Paleoecology and life history of Mastodonsaurus giganteus Item: Title: Conclusions Item: Title: Acknowledgments Item: Title: References Info: Title: Growing giants: ontogeny and life history of the temnospondyl Mastodonsaurus giganteus (Stereospondyli) from the Middle Triassic of Germany Author: Rainer R. Schoch Subject: The Middle Triassic capitosaur Mastodonsaurus giganteus was the largest temnospondyl and the dominating aquatic predator in many European freshwater to brackish ecosystems. It is represented by numerous size classes, which are described and analysed for the first time. The documented size range encompasses specimens between 12 15 mm and 1200 mm in skull length. Early growth stages are restricted to dentaries and interclavicles, whereas juveniles are represented by partial skulls, mandibles, and girdles. The smallest specimens already possessed diagnostic features of the taxon, and small juveniles also shared the dermal ornament with larger specimens. The heavy, disc-shaped intercentra were established early in the juvenile phase. Cranial proportions were remarkably conservative throughout ontogeny, with the orbits proportionately decreasing in size only very moderately, the postorbital skull becoming slightly longer and the occipital margin more concave in the largest forms. Analysis of frequency distributions of M. giganteus in different Lower Keuper deposits in southern Germany reflects habitat preferences in specific phases of its life cycle. The coal-bearing deposit at Gaildorf yielded unusually large specimens with relatively well-ossified appendicular skeletons. In the more common lake shore facies, only adult specimens are present. In turn, juveniles might have dwelled in calmer environments. Smaller lakes were apparently less attractive than larger or deeper water bodies that provided sufficient resources for several temnospondyls, and juvenile specimens have been identified from all of them. The diverse actinopterygian fish fauna provided prey for all growth stages of the large temnospondyl predators. 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It is represented by numerous size classes, which are described and analysed for the first time. The documented size range encompasses specimens between 12–15 mm and 1200 mm in skull length. Early growth stages are restricted to dentaries and interclavicles, whereas juveniles are represented by partial skulls, mandibles, and girdles. The smallest specimens already possessed diagnostic features of the taxon, and small juveniles also shared the dermal ornament with larger specimens. The heavy, disc-shaped intercentra were established early in the juvenile phase. Cranial proportions were remarkably conservative throughout ontogeny, with the orbits proportionately decreasing in size only very moderately, the postorbital skull becoming slightly longer and the occipital margin more concave in the largest forms. Analysis of frequency distributions of M. giganteus in different Lower Keuper deposits in southern Germany reflects habitat preferences in specific phases of its life cycle. The coal-bearing deposit at Gaildorf yielded unusually large specimens with relatively well-ossified appendicular skeletons. In the more common lake shore facies, only adult specimens are present. In turn, juveniles might have dwelled in calmer environments. Smaller lakes were apparently less attractive than larger or deeper water bodies that provided sufficient resources for several temnospondyls, and juvenile specimens have been identified from all of them. The diverse actinopterygian fish fauna provided prey for all growth stages of the large temnospondyl predators. Rainer R. Schoch et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Rainer R. Schoch Florian Witzmann Raphael Moreno Ralf Werneburg Eudald Mujal Capitosauria lower Keuper ontogeny Stereospondyli 2024-12-23T16:59:53+02:00 2024-12-23T17:00:03+02:00 2024-12-23T17:00:03+02:00 Adobe InDesign 20.0 (Windows) uuid:994cd52c-91fc-4e06-a969-dad0aa18e474 xmp.did:9F28E084BD8CE31184EC999A5D291C94 xmp.id:c2904904-9b89-c248-84ed-0fd874a6d102 proof:pdf xmp.iid:e6dbf39d-e504-8b42-83cc-8f3f0f26a9a9 xmp.did:4444FC51923DE811ADBCA6C78E600139 xmp.did:9F28E084BD8CE31184EC999A5D291C94 default converted from application/x-indesign to application/pdf Adobe InDesign 20.0 (Windows) / 2024-12-23T16:59:53+02:00 Adobe PDF Library 17.0 False Checksum: df16129f Type: CRC32 Checksum: c5987c0859c63a087232ca03834d8cd2 Type: MD5 Checksum: 1a4011876220bab8edb11dfcb6004bd0111d7123 Type: SHA-1 Checksum: 35de58d4e7051ec892dea099ca0abc2676d3273552d0b4c09fb52db9ffc5720d Type: SHA-256