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Title:           The trichome pattern diversity of Cardamine shares genetic mechanisms with Arabidopsis but differs in environmental drivers
Subject:         DOI: 10.1093/plphys/kiae213; Plant Physiology, 196, 4, 2024-04-12.; Abstract: Natural variation in trichome pattern (amount and distribution) is prominent among populations of many angiosperms. However, the degree of parallelism in the genetic mechanisms underlying this diversity and its environmental drivers in different species remain unclear. To address these questions, we analyzed the genomic and environmental bases of leaf trichome pattern diversity in Cardamine hirsuta, a relative of Arabidopsis (Arabidopsis thaliana). We characterized 123 wild accessions for their genomic diversity, leaf trichome patterns at different temperatures, and environmental adjustments. Nucleotide diversities and biogeographical distribution models identified two major genetic lineages with distinct demographic and adaptive histories. Additionally, C. hirsuta showed substantial variation in trichome pattern and plasticity to temperature. Trichome amount in C. hirsuta correlated positively with spring precipitation but negatively with temperature, which is opposite to climatic patterns in A. thaliana. Contrastingly, genetic analysis of C. hirsuta glabrous accessions indicated that, like for A. thaliana, glabrousness is caused by null mutations in ChGLABRA1 (ChGL1). Phenotypic genome-wide association studies (GWAS) further identified a ChGL1 haplogroup associated with low trichome density and ChGL1 expression. Therefore, a ChGL1 series of null and partial loss-of-function alleles accounts for the parallel evolution of leaf trichome pattern in C. hirsuta and A. thaliana. Finally, GWAS also detected other candidate genes (e.g. ChETC3, ChCLE17) that might affect trichome pattern. Accordingly, the evolution of this trait in C. hirsuta and A. thaliana shows partially conserved genetic mechanisms but is likely involved in adaptation to different environments.
Keywords:        
Author:          Alberto Fuster-Pons
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     Item: 
      Title: The trichome pattern diversity of Cardamine shares genetic mechanisms with Arabidopsis but differs in environmental drivers
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      Children: 
       Item: 
        Title: Introduction
        Destination: seckiae213-s0
       Item: 
        Title: Results
        Destination: seckiae213-s1
        Children: 
         Item: 
          Title: Genetic, geographic, and environmental structure of Cardamine nucleotide diversity
          Destination: seckiae213-s1.1
         Item: 
          Title: Ontogenic and genetic variation for leaf trichome pattern
          Destination: seckiae213-s1.2
         Item: 
          Title: Natural diversity for leaf trichome pattern at different temperatures
          Destination: seckiae213-s1.3
         Item: 
          Title: Geographic and climatic distribution of leaf trichome pattern diversity
          Destination: seckiae213-s1.4
         Item: 
          Title: GWAS of leaf trichome pattern and climatic variables
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         Item: 
          Title: Functional characterization of a ChGL1 allelic series affecting leaf trichome pattern
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       Item: 
        Title: Discussion
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        Children: 
         Item: 
          Title: The Iberian Peninsula as a highly diverse region for comparative analyses of Cardamine and Arabidopsis
          Destination: seckiae213-s2.1
         Item: 
          Title: Different environmental drivers shape the leaf trichome diversity of Cardamine and Arabidopsis
          Destination: seckiae213-s2.2
         Item: 
          Title: Conserved genetic mechanisms account for the trichome diversity of Cardamine and Arabidopsis
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       Item: 
        Title: Materials and methods
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        Children: 
         Item: 
          Title: Plant material and environmental data
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         Item: 
          Title: Growth conditions and phenotypic analyses
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         Item: 
          Title: Distribution modeling and environmental analyses
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         Item: 
          Title: Genome sequencing and genetic structure
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         Item: 
          Title: Phenotypic and environmental GWAS
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         Item: 
          Title: ChGL1 sequencing, phylogenetic, and expression analyses
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         Item: 
          Title: Other statistical analyses
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         Item: 
          Title: Accession numbers
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        Title: Acknowledgements
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        Title: Author contributions
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        Title: Supplementary data
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       Item: 
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    Title: The trichome pattern diversity of Cardamine shares genetic mechanisms with Arabidopsis but differs in environmental drivers
    Author: Alberto Fuster-Pons
    Subject: DOI: 10.1093/plphys/kiae213; Plant Physiology, 196, 4, 2024-04-12.; Abstract: Natural variation in trichome pattern (amount and distribution) is prominent among populations of many angiosperms. However, the degree of parallelism in the genetic mechanisms underlying this diversity and its environmental drivers in different species remain unclear. To address these questions, we analyzed the genomic and environmental bases of leaf trichome pattern diversity in Cardamine hirsuta, a relative of Arabidopsis (Arabidopsis thaliana). We characterized 123 wild accessions for their genomic diversity, leaf trichome patterns at different temperatures, and environmental adjustments. Nucleotide diversities and biogeographical distribution models identified two major genetic lineages with distinct demographic and adaptive histories. Additionally, C. hirsuta showed substantial variation in trichome pattern and plasticity to temperature. Trichome amount in C. hirsuta correlated positively with spring precipitation but negatively with temperature, which is opposite to climatic patterns in A. thaliana. Contrastingly, genetic analysis of C. hirsuta glabrous accessions indicated that, like for A. thaliana, glabrousness is caused by null mutations in ChGLABRA1 (ChGL1). Phenotypic genome-wide association studies (GWAS) further identified a ChGL1 haplogroup associated with low trichome density and ChGL1 expression. Therefore, a ChGL1 series of null and partial loss-of-function alleles accounts for the parallel evolution of leaf trichome pattern in C. hirsuta and A. thaliana. Finally, GWAS also detected other candidate genes (e.g. ChETC3, ChCLE17) that might affect trichome pattern. Accordingly, the evolution of this trait in C. hirsuta and A. thaliana shows partially conserved genetic mechanisms but is likely involved in adaptation to different environments.
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   Type: CRC32
  Checksum: 98b56152779314d32395859358baf2fb
   Type: MD5
  Checksum: 538853e4bb50a27f649a7afaa6f5b7df8fc8f23e
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   Type: SHA-256