808a6fb2e5c209a33021936f13c63513 Garrouste_etal_2025.pdf 9fc9ddfcd6931caf18f90162eb65a1b6d7cde32c Garrouste_etal_2025.pdf a1c0665b35dbe31efab3932e46c2d8895a76b43d780d72a74d4ae5a4e2c67ecb Garrouste_etal_2025.pdf Title: New insects from the Permian of Catalonia (Spain) predate the early diversification of Triassic clades adapted to arid environments (Grylloblattodea, Probnidae, Dictyoptera) Subject: Probnis sauvanyaensis sp. nov., the first representative of the ‘grylloblattodean’ family Probnidae from the Iberian Peninsula, is described from the middle to ?upper Permian outcrop of Sauvanyà in the Catalan Pyrenees. The family is otherwise represented by two additional species of Probnis from the ?middle Permian of the USA and by a single genus and species from the ?Lower Triassic of Ukraine. As is known from outcrops with rather extreme climates, the family may have survived the global warming of the Permian–Triassic crisis. We also describe a roachoid of the family Spiloblattidae from the same outcrop as the second representative of this family from the Iberian Peninsula. Author: Romain Garrouste Creator: Adobe InDesign 20.3 (Windows) Producer: Adobe PDF Library 17.0 CreationDate: Mon Jun 9 14:50:50 2025 CEST ModDate: Mon Jun 9 14:50:52 2025 CEST Custom Metadata: no Metadata Stream: yes Tagged: no UserProperties: no Suspects: no Form: none JavaScript: no Pages: 8 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 8163673 bytes Optimized: yes PDF version: 1.4 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- PZMVGO+TimesNewRomanPSMT CID TrueType Identity-H yes yes yes 160 0 MNZYQU+TimesNewRomanPS-ItalicMT CID TrueType Identity-H yes yes yes 170 0 SJZIWI+SymbolMT CID TrueType Identity-H yes yes yes 171 0 IZYFMC+NewsGothicStd Type 1C Custom yes yes yes 172 0 QGAWUM+FFMarkPro-Bold Type 1C WinAnsi yes yes yes 173 0 PKJLBO+NewsGothicStd-Oblique Type 1C WinAnsi yes yes yes 186 0 ZBPDQU+TimesNewRomanPSMT TrueType WinAnsi yes yes yes 166 0 KCCROY+NewsGothicMT TrueType WinAnsi yes yes yes 167 0 AVAEES+TimesNewRomanPS-ItalicMT TrueType WinAnsi yes yes yes 175 0 HQKUYE+TimesNewRomanPS-BoldMT TrueType WinAnsi yes yes yes 176 0 IZYFMC+NewsGothicStd Type 1C WinAnsi yes yes yes 108 0 IZYFMC+NewsGothicStd Type 1C WinAnsi yes yes yes 110 0 DSKRPM+TimesNewRomanPS-BoldItalicMT TrueType WinAnsi yes yes yes 114 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2025-06-11 02:07:46 CEST RepresentationInformation: Garrouste_etal_2025.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2025-06-10 12:16:12 CEST Size: 8163673 Format: PDF Version: 1.4 Status: Well-Formed and valid SignatureMatches: PDF-hul MIMEtype: application/pdf Profile: Linearized PDF PDFMetadata: Objects: 227 FreeObjects: 1 IncrementalUpdates: 1 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Language: en-US Outlines: Item: Title: New insects from the Permian of Catalonia (Spain) predate the early diversification of Triassic clades adapted to arid environments (Grylloblattodea, Probnidae, Dictyoptera) Item: Title: Abstract Item: Title: Introduction Children: Item: Title: Geological setting Item: Title: Materials and methods Children: Item: Title: Institutional abbreviations Item: Title: Systematic paleontology Children: Item: Title: Class INSECTA Linnæus, 1758 Item: Title: Genus Probnis Sellards, 1909 Item: Title: Probnis sauvanyaensis sp. nov. Item: Title: Superorder ‘HOLOPANDICTYOPTERA Kluge, 2010’ (= total group of extant Dictyoptera Leach, 1815) Item: Title: Family SPILOBLATTINIDAE Handlirsch, 1906 Item: Title: Discussion and conclusion Item: Title: Acknowledgments Item: Title: References Info: Title: New insects from the Permian of Catalonia (Spain) predate the early diversification of Triassic clades adapted to arid environments (Grylloblattodea, Probnidae, Dictyoptera) Author: Romain Garrouste Subject: Probnis sauvanyaensis sp. nov., the first representative of the ‘grylloblattodean’ family Probnidae from the Iberian Peninsula, is described from the middle to ?upper Permian outcrop of Sauvanyà in the Catalan Pyrenees. The family is otherwise represented by two additional species of Probnis from the ?middle Permian of the USA and by a single genus and species from the ?Lower Triassic of Ukraine. As is known from outcrops with rather extreme climates, the family may have survived the global warming of the Permian Triassic crisis. We also describe a roachoid of the family Spiloblattidae from the same outcrop as the second representative of this family from the Iberian Peninsula. 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The family is otherwise represented by two additional species of Probnis from the ?middle Permian of the USA and by a single genus and species from the ?Lower Triassic of Ukraine. As is known from outcrops with rather extreme climates, the family may have survived the global warming of the Permian–Triassic crisis. We also describe a roachoid of the family Spiloblattidae from the same outcrop as the second representative of this family from the Iberian Peninsula. Romain Garrouste et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 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