d8da8147d51d132d1a41b89a5bb08748 diversity-17-00631.pdf 6cbcf4647c775b85c3355976c6bd67f14f6d6392 diversity-17-00631.pdf 531a1846c4319fc08679298530dd501094bc97a633e4c82aa61d4884954cee85 diversity-17-00631.pdf Title: An Integrated Paleoenvironmental Reconstruction of the Early Pleistocene Hominin-Bearing Site of Dursunlu (Türkiye) Subject: The fossiliferous Dursunlu Lignite Quarry (DLQ) is highlighted prominently in the archeological and paleontological literature because of the study of Pleistocene fauna and lithic artifacts, being considered the oldest Paleolithic site in Türkiye. Although the fauna and flora assemblage from DLQ are reasonably well known, taxonomic studies devoted to some groups, such as ostracods, mollusks, reptiles, and amphibians, have never been carried out. Here, we describe, illustrate, and study the taxonomic composition and ecological implications of the unpublished material of said groups, together with the aquatic plants and fish, recovered from six samples taken from the palustrine and peat bog facies of the sedimentary sequence. In addition, the recovered charophytes and cyprinids refine our taxonomical knowledge of both aquatic plants and fish. Our results concur with previous paleoenvironmental inferences based on flora and fauna composition—with DLQ representing a very shallow eutrophic lake with a dense palustrine vegetation belt during the cold (glacial) stage of the late Early Pleistocene—as well as highlight the study of all available groups as pivotal for better understanding the paleolake biota. We further conclude that the wetland areas of Dursunlu and surrounding steppe areas appear to have been an excellent environment for sporadic settlement of hominins during the Early Pleistocene, given the availability of food resources and easy access to water. Keywords: Dursunlu paleolake; Günz glaciation; paleoecology; hominins sporadic settlement; Central Anatolia Author: Àngel H. Luján, Václav Paclík, Elvan Demirci, Andrea Villa, Thomas A. Neubauer, Alaettin Tuncer, Martin Ivanov, Àngel Blanco-Lapaz, Kelly Ann Vega-Pagán and Josep Sanjuan Creator: LaTeX with hyperref Producer: pdfTeX-1.40.25; modified using OpenPDF 1.4.2 CreationDate: Mon Sep 8 11:23:17 2025 CEST ModDate: Mon Sep 8 13:12:12 2025 CEST Custom Metadata: yes Metadata Stream: no Tagged: no UserProperties: no Suspects: no Form: none JavaScript: no Pages: 31 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 23510990 bytes Optimized: no PDF version: 1.5 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- PGPDYS+VnURWPalladioL-Bold Type 1 Custom yes yes yes 101 0 MJOOAC+URWPalladioL-Roma Type 1 Custom yes yes yes 102 0 PSHMZA+VnURWPalladioL Type 1 Custom yes yes yes 103 0 CCRHYS+URWPalladioL-Ital Type 1 Custom yes yes yes 104 0 XTSVRM+URWPalladioL-Bold Type 1 Custom yes yes yes 105 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 134 0 TRVXNW+CMSY10 Type 1 Builtin yes yes yes 184 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 198 0 BDENFB+PalatinoLinotype,Italic TrueType WinAnsi yes yes no 372 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 373 0 OVNEXW+EURM10 Type 1 Builtin yes yes yes 412 0 BDENFB+PalatinoLinotype,Italic TrueType WinAnsi yes yes no 379 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 380 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 407 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 426 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 458 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 528 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 533 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 548 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 754 0 BDENDP+PalatinoLinotype TrueType WinAnsi yes yes no 793 0 BDENFB+PalatinoLinotype,Italic TrueType WinAnsi yes yes no 794 0 ESSRQF+VnURWPalladioL-Italic Type 1 Custom yes yes yes 895 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2025-09-17 03:09:49 CEST RepresentationInformation: diversity-17-00631.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2025-09-16 09:34:11 CEST Size: 23510990 Format: PDF Version: 1.5 Status: Well-Formed and valid SignatureMatches: PDF-hul MIMEtype: application/pdf PDFMetadata: Objects: 1010 FreeObjects: 1 IncrementalUpdates: 0 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: true CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Outlines: Item: Title: Introduction Children: Item: Title: The Hominin-Bearing Site of Dursunlu Lignite Quarry Children: Item: Title: Historical Background Item: Title: Chronological Background and Aims Item: Title: Geographical and Geological Setting Item: Title: Material and Methods Item: Title: Results Item: Title: Discussion Children: Item: Title: Paleoenvironmental Reconstruction of Dursunlu Paleolake Item: Title: Potential Resources for Homo During the Early Pleistocene Item: Title: Conclusions Item: Title: References Info: Title: An Integrated Paleoenvironmental Reconstruction of the Early Pleistocene Hominin-Bearing Site of Dursunlu (Türkiye) Author: Àngel H. Luján, Václav Paclík, Elvan Demirci, Andrea Villa, Thomas A. Neubauer, Alaettin Tuncer, Martin Ivanov, Àngel Blanco-Lapaz, Kelly Ann Vega-Pagán and Josep Sanjuan Subject: The fossiliferous Dursunlu Lignite Quarry (DLQ) is highlighted prominently in the archeological and paleontological literature because of the study of Pleistocene fauna and lithic artifacts, being considered the oldest Paleolithic site in Türkiye. Although the fauna and flora assemblage from DLQ are reasonably well known, taxonomic studies devoted to some groups, such as ostracods, mollusks, reptiles, and amphibians, have never been carried out. Here, we describe, illustrate, and study the taxonomic composition and ecological implications of the unpublished material of said groups, together with the aquatic plants and fish, recovered from six samples taken from the palustrine and peat bog facies of the sedimentary sequence. In addition, the recovered charophytes and cyprinids refine our taxonomical knowledge of both aquatic plants and fish. Our results concur with previous paleoenvironmental inferences based on flora and fauna composition—with DLQ representing a very shallow eutrophic lake with a dense palustrine vegetation belt during the cold (glacial) stage of the late Early Pleistocene—as well as highlight the study of all available groups as pivotal for better understanding the paleolake biota. We further conclude that the wetland areas of Dursunlu and surrounding steppe areas appear to have been an excellent environment for sporadic settlement of hominins during the Early Pleistocene, given the availability of food resources and easy access to water. 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