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referenced object 'WEB_COPLBI' cannot be found ID: PDF-HUL-149 ErrorMessage: edu.harvard.hul.ois.jhove.module.pdf.PdfInvalidException: Invalid indirect destination - referenced object 'WEB_COPLBI' cannot be found ID: PDF-HUL-122 Offset: 2484060 MIMEtype: application/pdf Profile: Linearized PDF, Tagged PDF PDFMetadata: Objects: 1855 FreeObjects: 1 IncrementalUpdates: 1 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: true NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseOutlines Language: en Outlines: Item: Title: Non-coding regulation in seasonal flowering control   Insights from FLC Destination: titlink1 Children: Item: Title: Introduction Destination: flink1 Item: Title: Molecular basis of vernalization: cis-regulatory elements and long non-coding RNAs Destination: flink2 Item: Title: Non-coding polymorphisms modulate FLC expression dynamics Destination: flink3 Item: Title: Vernalization variation drives divergence in plant life history strategies Destination: flink4 Item: Title: Current challenges and future perspectives Destination: flink5 Item: Title: Author contributions Destination: flink6 Item: Title: Declaration of competing interest Destination: creditlink2 Item: Title: Acknowledgements Destination: aclink3 Item: Title: Data availability Destination: datalink4 Item: Title: References Destination: blink1 Info: Title: Non-coding regulation in seasonal flowering control - Insights from FLC Author: Mélanie Ormancey Subject: Current Opinion in Plant Biology, 89 (2026) 102831. doi:10.1016/j.pbi.2025.102831 Creator: Elsevier Producer: PDFlib+PDI 10.0.2p2 (C++/Win64) CreationDate: Fri Dec 12 12:15:49 CET 2025 ModDate: Fri Dec 12 12:20:07 CET 2025 ID: 0x4bb47cdc1656dec0765692216aac6eae, 0xbce4bc6739687d47bdeca9af77c9cc15 Filters: FilterPipeline: FlateDecode Fonts: Type0: Font: BaseFont: GlyphLessFont Encoding: Identity-H ToUnicode: true Font: BaseFont: HPYGKY+Helvetica Encoding: Identity-H ToUnicode: true Type1: Font: BaseFont: OURBKI+HelveticaLTStd-Bold FontSubset: true FirstChar: 1 LastChar: 73 FontDescriptor: FontName: OURBKI+HelveticaLTStd-Bold Flags: Nonsymbolic, ForceBold FontBBox: -170, -228, 1003, 962 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: DJKDIV+ESSTIX-Five FontSubset: true FirstChar: 33 LastChar: 116 FontDescriptor: FontName: DJKDIV+ESSTIX-Five Flags: Symbolic FontBBox: -50, -180, 1000, 720 FontFile3: true ToUnicode: true Font: BaseFont: YVOEHY+Caslon540BT-Roman FontSubset: true FirstChar: 33 LastChar: 251 FontDescriptor: FontName: YVOEHY+Caslon540BT-Roman Flags: Symbolic FontBBox: -50, -180, 1000, 720 FontFile3: true ToUnicode: true Font: BaseFont: KOGBIL+HelveticaLTStd-BoldObl FontSubset: true FirstChar: 1 LastChar: 17 FontDescriptor: FontName: KOGBIL+HelveticaLTStd-BoldObl Flags: Nonsymbolic, Italic, ForceBold FontBBox: -174, -228, 1114, 962 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: QLRHEX+CM_Maths_Symbols FontSubset: true FirstChar: 1 LastChar: 138 FontDescriptor: FontName: QLRHEX+CM_Maths_Symbols Flags: Symbolic FontBBox: -50, -180, 1000, 720 FontFile3: true ToUnicode: true Font: BaseFont: BKIITU+Caslon540BT-Italic FontSubset: true FirstChar: 32 LastChar: 251 FontDescriptor: FontName: BKIITU+Caslon540BT-Italic Flags: Symbolic FontBBox: -50, -180, 1000, 720 FontFile3: true ToUnicode: true Font: BaseFont: PFJEEC+HelveticaLTStd-Roman FontSubset: true FirstChar: 1 LastChar: 83 FontDescriptor: FontName: PFJEEC+HelveticaLTStd-Roman Flags: Nonsymbolic FontBBox: -166, -225, 1000, 931 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: RWJVUX+VAGRoundedStd-Bold FontSubset: true FirstChar: 1 LastChar: 8 FontDescriptor: FontName: RWJVUX+VAGRoundedStd-Bold Flags: Nonsymbolic, ForceBold FontBBox: -151, -310, 1060, 939 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true Font: BaseFont: YHNHDF+HelveticaLTStd-Obl FontSubset: true FirstChar: 1 LastChar: 44 FontDescriptor: FontName: YHNHDF+HelveticaLTStd-Obl Flags: Nonsymbolic, Italic FontBBox: -170, -225, 1116, 931 FontFile3: true EncodingDictionary: BaseEncoding: WinAnsiEncoding Differences: true ToUnicode: true CIDFontType2: Font: BaseFont: GlyphLessFont CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: GlyphLessFont Flags: FixedPitch, Symbolic FontBBox: 0, 0, 500, 1000 FontFile2: true Font: BaseFont: HPYGKY+Helvetica CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: HPYGKY+Helvetica Flags: Nonsymbolic FontBBox: 0, -225, 1000, 931 FontFile2: true XMP: application/pdf doi:10.1016/j.pbi.2025.102831 Non-coding regulation in seasonal flowering control - Insights from FLC Mélanie Ormancey Julia I. Qüesta FLOWERING LOCUS C Seasonality Vernalization Cold temperature Non-coding polymorphisms Long non-coding RNAs Life history Flowering time Current Opinion in Plant Biology, 89 (2026) 102831. doi:10.1016/j.pbi.2025.102831 The Author(s) journal Current Opinion in Plant Biology © 2025 The Author(s). 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