6acfcbb552cfdb7b445faad57291e8ac pnas_a2025m12v123n1p2519908122.pdf 987439888acbdcac55bd74b4c4ed18c82d15543d pnas_a2025m12v123n1p2519908122.pdf eb630bf68e7b311e799ff75f7e3312dec4f7639f2dbf2deb79405c02b703dd02 pnas_a2025m12v123n1p2519908122.pdf Custom Metadata: no Metadata Stream: no Tagged: no UserProperties: no Suspects: no Form: none JavaScript: no Pages: 17 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 466200 bytes Optimized: no PDF version: 1.7 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- OPKMFX+LibertinusSerif-Regular-Identity-H CID Type 0C Identity-H yes yes yes 60 0 WNVUHC+LibertinusSerif-Bold-Identity-H CID Type 0C Identity-H yes yes yes 61 0 WOTGTX+LibertinusSerif-Italic-Identity-H CID Type 0C Identity-H yes yes yes 62 0 AAAAAH+ArialMT CID TrueType Identity-H yes yes yes 73 0 AAAAAC+Calibri TrueType WinAnsi yes yes yes 74 0 AAAAAE+Arial-BoldMT TrueType WinAnsi yes yes yes 75 0 AAAAAG+ArialMT TrueType WinAnsi yes yes yes 76 0 AAAAAJ+Cambria TrueType WinAnsi yes yes yes 78 0 AAAAAL+Arial-ItalicMT TrueType WinAnsi yes yes yes 79 0 AAAAAN+Arial-BoldItalicMT TrueType WinAnsi yes yes yes 80 0 AAAAAO+Arial-ItalicMT CID TrueType Identity-H yes yes yes 81 0 AAAAAP+ArialUnicodeMS CID TrueType Identity-H yes yes yes 82 0 AAAAAR+ArialUnicodeMS TrueType WinAnsi yes yes yes 83 0 AAAAAS+CambriaMath CID TrueType Identity-H yes yes yes 84 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2026-01-10 04:49:06 CET RepresentationInformation: pnas_a2025m12v123n1p2519908122.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2026-01-09 12:43:14 CET Size: 466200 Format: PDF Version: 1.7 Status: Well-Formed and valid SignatureMatches: PDF-hul MIMEtype: application/pdf PDFMetadata: Objects: 152 FreeObjects: 1 IncrementalUpdates: 0 DocumentCatalog: PageLayout: SinglePage PageMode: UseNone ID: 0xe28603e7ce76dc82743819abcbf93b2d, 0xe28603e7ce76dc82743819abcbf93b2d Filters: FilterPipeline: FlateDecode FilterPipeline: DCTDecode Images: Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 228 ImageHeight: 127 BitsPerSample: 8 BitsPerSampleUnit: integer ColorSpace: ICCBased Interpolate: false Image: NisoImageMetadata: FormatName: image/png ImageWidth: 296 ImageHeight: 104 BitsPerSample: 8 BitsPerSampleUnit: integer Filter: FlateDecode Interpolate: true Fonts: Type0: Font: BaseFont: AAAAAO+Arial-ItalicMT Encoding: Identity-H ToUnicode: true Font: BaseFont: AAAAAP+ArialUnicodeMS Encoding: Identity-H ToUnicode: true Font: BaseFont: AAAAAS+CambriaMath Encoding: Identity-H ToUnicode: true Font: BaseFont: AAAAAH+ArialMT Encoding: Identity-H ToUnicode: true Font: BaseFont: OPKMFX+LibertinusSerif-Regular-Identity-H Encoding: Identity-H ToUnicode: true Font: BaseFont: WNVUHC+LibertinusSerif-Bold-Identity-H Encoding: Identity-H ToUnicode: true Font: BaseFont: WOTGTX+LibertinusSerif-Italic-Identity-H Encoding: Identity-H ToUnicode: true TrueType: Font: BaseFont: AAAAAN+Arial-BoldItalicMT FontSubset: true FirstChar: 33 LastChar: 39 FontDescriptor: FontName: AAAAAN+Arial-BoldItalicMT Flags: Symbolic, Italic FontBBox: -560, -376, 1390, 1018 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAR+ArialUnicodeMS FontSubset: true FirstChar: 33 LastChar: 33 FontDescriptor: FontName: AAAAAR+ArialUnicodeMS Flags: Symbolic FontBBox: -1011, -330, 2260, 1078 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAC+Calibri FontSubset: true FirstChar: 33 LastChar: 44 FontDescriptor: FontName: AAAAAC+Calibri Flags: Symbolic FontBBox: -503, -313, 1240, 1026 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAE+Arial-BoldMT FontSubset: true FirstChar: 33 LastChar: 89 FontDescriptor: FontName: AAAAAE+Arial-BoldMT Flags: Symbolic FontBBox: -628, -376, 2000, 1056 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAG+ArialMT FontSubset: true FirstChar: 33 LastChar: 121 FontDescriptor: FontName: AAAAAG+ArialMT Flags: Symbolic FontBBox: -665, -325, 2000, 1039 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAJ+Cambria FontSubset: true FirstChar: 33 LastChar: 33 FontDescriptor: FontName: AAAAAJ+Cambria Flags: Symbolic FontBBox: -1475, -2464, 2867, 3117 FontFile2: true ToUnicode: true Font: BaseFont: AAAAAL+Arial-ItalicMT FontSubset: true FirstChar: 33 LastChar: 76 FontDescriptor: FontName: AAAAAL+Arial-ItalicMT Flags: Symbolic, Italic FontBBox: -517, -325, 1359, 997 FontFile2: true ToUnicode: true CIDFontType0: Font: BaseFont: OPKMFX+LibertinusSerif-Regular CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: OPKMFX+LibertinusSerif-Regular Flags: Serif, Symbolic, SmallCap FontBBox: -6275, -256, 6171, 1125 FontFile3: true Font: BaseFont: WNVUHC+LibertinusSerif-Bold CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: WNVUHC+LibertinusSerif-Bold Flags: Serif, Symbolic, SmallCap FontBBox: -6275, -328, 6171, 1014 FontFile3: true Font: BaseFont: WOTGTX+LibertinusSerif-Italic CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: WOTGTX+LibertinusSerif-Italic Flags: Serif, Symbolic, Italic, SmallCap FontBBox: -6275, -312, 6171, 928 FontFile3: true CIDFontType2: Font: BaseFont: AAAAAO+Arial-ItalicMT CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: AAAAAO+Arial-ItalicMT Flags: Symbolic, Italic FontBBox: -517, -325, 1359, 997 FontFile2: true Font: BaseFont: AAAAAH+ArialMT CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: AAAAAH+ArialMT Flags: Symbolic FontBBox: -665, -325, 2000, 1039 FontFile2: true Font: BaseFont: AAAAAP+ArialUnicodeMS CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: AAAAAP+ArialUnicodeMS Flags: Symbolic FontBBox: -1011, -330, 2260, 1078 FontFile2: true Font: BaseFont: AAAAAS+CambriaMath CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: AAAAAS+CambriaMath Flags: Symbolic FontBBox: -1475, -2464, 2867, 3117 FontFile2: true Pages: Page: Sequence: 1 Annotations: Annotation: Subtype: Link Rect: 194, 496, 373, 503 Flags: Print Annotation: Subtype: Link Rect: 241, 462, 391, 469 Flags: Print Page: Sequence: 2 Annotations: Annotation: Subtype: Link Rect: 303, 192, 380, 204 Flags: Print AppearanceDictionary: true Annotation: Subtype: Link Rect: 136, 169, 234, 181 Flags: Print AppearanceDictionary: true Page: Sequence: 3 Page: Sequence: 4 Page: Sequence: 5 Page: Sequence: 6 Page: Sequence: 7 Page: Sequence: 8 Page: Sequence: 9 Page: Sequence: 10 Page: Sequence: 11 Page: Sequence: 12 Page: Sequence: 13 Page: Sequence: 14 Page: Sequence: 15 Page: Sequence: 16 Page: Sequence: 17 Checksum: a292577d Type: CRC32 Checksum: 6acfcbb552cfdb7b445faad57291e8ac Type: MD5 Checksum: 987439888acbdcac55bd74b4c4ed18c82d15543d Type: SHA-1 Checksum: eb630bf68e7b311e799ff75f7e3312dec4f7639f2dbf2deb79405c02b703dd02 Type: SHA-256