ae3f361191cd1e8cbe27c8012183f131 foss-rec_article_183059_en_1.pdf 1a0709f3cfdb2ba9762b36f9652603486e5cf666 foss-rec_article_183059_en_1.pdf 381de5fcf9a9bd0074037930103ad88bf63b87cc9565bd38104cac39449fd33d foss-rec_article_183059_en_1.pdf Title: First anguid record (Anguidae, Squamata) from the Late Miocene Konservat-Lagerstätte Libros (Teruel, Spain) and its palaeoenvironmental implications Subject: For the first time, fossil evidence of anguine material is reported from the middle–late Tortonian Konservat-Lagerstätte locality of Libros (Teruel, Spain). So far, only mammals, birds, snakes, amphibians, insects and flora in excellent conditions of preservation were described from the lacustrine sediments of this site. The size and fragility of the new fossil, partially embedded in the sediment, required the application of non-invasive techniques for their study. Micro-CT scans were conducted, resulting in satisfactory three-dimensional models. The skeleton is partially articulated and preserves the nasal bone, dozens of vertebrae, eleven ribs and hundreds of osteoderms. Amongst them, the two best-preserved trunk vertebrae and one caudal vertebra, along with osteoderm dermal sculpturing, suggest attribution to the genus Pseudopus (Family Anguidae). This new fossil record not only provides information from a global palaeobiogeographical perspective, but also constitutes a valuable contribution to the understanding of the ecosystems surrounding the Libros palaeolake. Author: Adriana Linares-Martín Creator: Adobe InDesign 21.4 (Windows) Producer: Adobe PDF Library 18.0 CreationDate: Fri Jun 19 14:32:51 2026 CEST ModDate: Fri Jun 19 14:32:54 2026 CEST Custom Metadata: no Metadata Stream: yes Tagged: yes UserProperties: no Suspects: no Form: none JavaScript: no Pages: 18 Encrypted: no Page size: 595.276 x 841.89 pts (A4) Page rot: 0 File size: 6446853 bytes Optimized: yes PDF version: 1.4 name type encoding emb sub uni object ID ------------------------------------ ----------------- ---------------- --- --- --- --------- FGJMHJ+TimesNewRomanPSMT CID TrueType Identity-H yes yes yes 1102 0 ZFFXBV+TimesNewRomanPS-ItalicMT CID TrueType Identity-H yes yes yes 1103 0 NVEYPT+SymbolMT CID TrueType Identity-H yes yes yes 1122 0 GAUNVH+NewsGothicStd Type 1C Custom yes yes yes 1104 0 EFTWTL+FFMarkPro-Bold Type 1C WinAnsi yes yes yes 1123 0 OHREDR+NewsGothicStd-Oblique Type 1C WinAnsi yes yes yes 1129 0 MBTXBV+TimesNewRomanPSMT TrueType WinAnsi yes yes yes 1106 0 SXTHHJ+TimesNewRomanPS-ItalicMT TrueType WinAnsi yes yes yes 1107 0 GAUNVH+NewsGothicMT TrueType WinAnsi yes yes yes 1108 0 EFTWTL+TimesNewRomanPS-BoldMT TrueType WinAnsi yes yes yes 1126 0 GAUNVH+NewsGothicStd Type 1C WinAnsi yes yes yes 62 0 GAUNVH+NewsGothicStd Type 1C WinAnsi yes yes yes 65 0 Jhove (Rel. 1.28.0, 2023-05-18) Date: 2026-06-23 03:57:03 CEST RepresentationInformation: foss-rec_article_183059_en_1.pdf ReportingModule: PDF-hul, Rel. 1.12.4 (2023-03-16) LastModified: 2026-06-22 15:16:24 CEST Size: 6446853 Format: PDF Version: 1.4 Status: Well-Formed, but not valid SignatureMatches: PDF-hul ErrorMessage: Annotation object is not a dictionary ID: PDF-HUL-114 Offset: 5382 MIMEtype: application/pdf Profile: Linearized PDF PDFMetadata: Objects: 1167 FreeObjects: 962 IncrementalUpdates: 1 DocumentCatalog: ViewerPreferences: HideToolbar: false HideMenubar: false HideWindowUI: false FitWindow: false CenterWindow: false DisplayDocTitle: false NonFullScreenPageMode: UseNone Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Language: en-US Outlines: Item: Title: First anguid record (Anguidae, Squamata) from the Late Miocene Konservat-Lagerstätte Libros (Teruel, Spain) and its palaeoenvironmental implications Item: Title: Abstract Item: Title: Introduction Children: Item: Title: Geological context Item: Title: Material and methods Children: Item: Title: X-ray microtomography Item: Title: Specimens examined Item: Title: Data resources Item: Title: Results Children: Item: Title: Systematic Palaeontology Item: Title: Anatomical description using scanner technologies Item: Title: Taphonomic inferences Item: Title: Discussion Children: Item: Title: Taxonomic identification of the libros material Item: Title: Palaeobiogeography of genus Pseudopus during the Miocene and Pliocene Item: Title: Contribution to the palaeoenvironment of Libros site Item: Title: Conclusions Item: Title: Acknowledgements Item: Title: References Item: Title: Additional information Item: Title: Supplementary material 1 Children: Item: Title: Supplementary taxa Item: Title: Supplementary material 2 Children: Item: Title: Supplementary measurements Info: Title: First anguid record (Anguidae, Squamata) from the Late Miocene Konservat-Lagerstätte Libros (Teruel, Spain) and its palaeoenvironmental implications Author: Adriana Linares-Martín Subject: For the first time, fossil evidence of anguine material is reported from the middle late Tortonian Konservat-Lagerstätte locality of Libros (Teruel, Spain). So far, only mammals, birds, snakes, amphibians, insects and flora in excellent conditions of preservation were described from the lacustrine sediments of this site. The size and fragility of the new fossil, partially embedded in the sediment, required the application of non-invasive techniques for their study. Micro-CT scans were conducted, resulting in satisfactory three-dimensional models. The skeleton is partially articulated and preserves the nasal bone, dozens of vertebrae, eleven ribs and hundreds of osteoderms. Amongst them, the two best-preserved trunk vertebrae and one caudal vertebra, along with osteoderm dermal sculpturing, suggest attribution to the genus Pseudopus (Family Anguidae). This new fossil record not only provides information from a global palaeobiogeographical perspective, but also constitutes a valuable contribution to the understanding of the ecosystems surrounding the Libros palaeolake. 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So far, only mammals, birds, snakes, amphibians, insects and flora in excellent conditions of preservation were described from the lacustrine sediments of this site. The size and fragility of the new fossil, partially embedded in the sediment, required the application of non-invasive techniques for their study. Micro-CT scans were conducted, resulting in satisfactory three-dimensional models. The skeleton is partially articulated and preserves the nasal bone, dozens of vertebrae, eleven ribs and hundreds of osteoderms. Amongst them, the two best-preserved trunk vertebrae and one caudal vertebra, along with osteoderm dermal sculpturing, suggest attribution to the genus Pseudopus (Family Anguidae). This new fossil record not only provides information from a global palaeobiogeographical perspective, but also constitutes a valuable contribution to the understanding of the ecosystems surrounding the Libros palaeolake. Adriana Linares-Martín et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Adriana Linares-Martín Tomáš Přikryl Andrea Villa Massimo Delfino Bruno Gómez De Soler Hugues-Alexandre Blain Computed tomography Europe limbless lizard Pseudopus 2026-06-19T15:32:51+03:00 2026-06-19T15:32:54+03:00 2026-06-19T15:32:54+03:00 Adobe InDesign 21.4 (Windows) uuid:ef159baa-ba9c-4093-a3d5-2fea99162a2c xmp.did:9F28E084BD8CE31184EC999A5D291C94 xmp.id:e15b6f26-731c-314d-b48e-51524cc0582f proof:pdf xmp.iid:17357a9c-e43c-474c-afd9-25902607b221 xmp.did:4444FC51923DE811ADBCA6C78E600139 xmp.did:9F28E084BD8CE31184EC999A5D291C94 default converted from application/x-indesign to application/pdf Adobe InDesign 21.4 (Windows) / 2026-06-19T15:32:51+03:00 Adobe PDF Library 18.0 False Checksum: d7c75ef3 Type: CRC32 Checksum: ae3f361191cd1e8cbe27c8012183f131 Type: MD5 Checksum: 1a0709f3cfdb2ba9762b36f9652603486e5cf666 Type: SHA-1 Checksum: 381de5fcf9a9bd0074037930103ad88bf63b87cc9565bd38104cac39449fd33d Type: SHA-256