d939d655a09bba362a96541cd234eb71 SJP-145-267_article-183361_en_1.pdf dec7a3ce6f09cdf36c93a654b1a74fcdbe175006 SJP-145-267_article-183361_en_1.pdf 8ab7e5a3ed243638f6f925c2ce6cfc83af0067fff3b70e6d3dc0c1b019dc0046 SJP-145-267_article-183361_en_1.pdf Title: The suid assemblage from the Vallesian locality of Can Llobateres 1 (Vallès-Penedès Basin, NE Iberian Peninsula) revisited Subject: The suid assemblages from Europe experienced considerable turnover during the Vallesian (MN9-MN10, early Late Miocene), ultimately leading to a marked decline in diversity during the Turolian. To provide further insight into the timing and paleoecological context of this Vallesian turnover in the suid assemblages, here we revisit the taxonomic composition of the suids from the Late Miocene (9.8 Ma) fossil locality of Can Llobateres 1 (CLL1; Vallès-Penedès Basin, NE Iberian Peninsula), with emphasis on species other than the small tetraconodontine Parachleuastochoerus crusafonti (for which CLL1 is the type locality). Although CLL1 is one of the most thoroughly sampled Vallesian sites from Europe, a recent revision of the faunal list evinced that the occurrence of multiple large mammals is based on fossil remains that were described many decades ago or that have never been published, leading to potentially misleading or, at least, insufficiently substantiated attributions. According to published accounts, three additional suid species are recorded at CLL1, being much scarcer than Pa. crusafonti: the listriodontine Listriodon splendens, the suine Propotamochoerus palaeochoerus, and a large tetraconodontine variously assigned to Versoporcus steinheimensis or ‘Parachleuastochoerus’ valentini. Our revision of the available remains, including unpublished material recovered from the 1980s onward, confirms the presence of the two former species and the assignment of the large tetraconodontine to ‘Pa.’ valentini. On biochronological grounds, the association of these taxa is characteristic of the early Vallesian, with the record of L. splendens representing the last well-dated occurrence of this species. From a paleoenvironmental viewpoint, the suid assemblage from CLL1 is consistent with previous inferences indicating the presence of a densely forested and humid environment with more open habitats nearby. Author: Sara G. 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Direction: L2R ViewArea: CropBox ViewClip: CropBox PrintArea: CropBox PageClip: CropBox PageLayout: SinglePage PageMode: UseNone Language: en-US Outlines: Item: Title: The suid assemblage from the Vallesian locality of Can Llobateres 1 (Vallès-Penedès Basin, NE Iberian Peninsula) revisited Item: Title: Abstract Item: Title: Introduction Children: Item: Title: Taxonomic controversies about European tetraconodontines Item: Title: The suid assemblage from Can Llobateres 1 Item: Title: Materials and methods Children: Item: Title: Studied sample Item: Title: Comparative sample Item: Title: Dental terminology and measurements Item: Title: Abbreviations Item: Title: Systematic Paleontology Item: Title: Listriodon splendens Von Meyer, 1846 Item: Title: Subfamily Tetraconodontinae Lydekker, 1876 Item: Title: ‘Parachleuastochoerus’ valentini (Filhol, 1882) Item: Title: Subfamily Suinae Gray, 1821 Item: Title: Propotamochoerus palaeochoerus (Kaup, 1833) Item: Title: Discussion and conclusions Item: Title: Acknowledgments Item: Title: Additional information Item: Title: References Item: Title: Supplementary material 1 Children: Item: Title: Supplementary Information Filters: FilterPipeline: FlateDecode FilterPipeline: DCTDecode Images: Image: NisoImageMetadata: FormatName: image/png ImageWidth: 1994 ImageHeight: 2837 ColorSpace: CMYK BitsPerSample: 8 BitsPerSampleUnit: integer Filter: FlateDecode Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 2001 ImageHeight: 2272 ColorSpace: CMYK BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 2005 ImageHeight: 733 ColorSpace: CMYK BitsPerSample: 8 BitsPerSampleUnit: integer Intent: RelativeColorimetric Name: X Image: NisoImageMetadata: FormatName: image/jpg CompressionScheme: JPEG ImageWidth: 2001 ImageHeight: 2214 ColorSpace: CMYK BitsPerSample: 8 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Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: MGLQIK+TimesNewRomanPS-BoldMT Flags: Serif, Symbolic FontBBox: -558, -307, 2000, 1026 FontFile2: true Font: BaseFont: SHPFOY+SymbolMT CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: SHPFOY+SymbolMT Flags: Symbolic FontBBox: 0, -220, 1113, 1005 FontFile2: true Font: BaseFont: SJQKOY+TimesNewRomanPSMT CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: SJQKOY+TimesNewRomanPSMT Flags: Serif, Symbolic FontBBox: -568, -307, 2046, 1040 FontFile2: true Font: BaseFont: EEOZAA+TimesNewRomanPS-ItalicMT CIDSystemInfo: Registry: Adobe Registry: Identity Supplement: 0 FontDescriptor: FontName: EEOZAA+TimesNewRomanPS-ItalicMT Flags: Serif, Symbolic, Italic FontBBox: -498, -307, 1353, 1023 FontFile2: true XMP: application/pdf The suid assemblage from the Vallesian locality of Can Llobateres 1 (Vallès-Penedès Basin, NE Iberian Peninsula) revisited The suid assemblages from Europe experienced considerable turnover during the Vallesian (MN9-MN10, early Late Miocene), ultimately leading to a marked decline in diversity during the Turolian. To provide further insight into the timing and paleoecological context of this Vallesian turnover in the suid assemblages, here we revisit the taxonomic composition of the suids from the Late Miocene (9.8 Ma) fossil locality of Can Llobateres 1 (CLL1; Vallès-Penedès Basin, NE Iberian Peninsula), with emphasis on species other than the small tetraconodontine Parachleuastochoerus crusafonti (for which CLL1 is the type locality). Although CLL1 is one of the most thoroughly sampled Vallesian sites from Europe, a recent revision of the faunal list evinced that the occurrence of multiple large mammals is based on fossil remains that were described many decades ago or that have never been published, leading to potentially misleading or, at least, insufficiently substantiated attributions. According to published accounts, three additional suid species are recorded at CLL1, being much scarcer than Pa. crusafonti: the listriodontine Listriodon splendens, the suine Propotamochoerus palaeochoerus, and a large tetraconodontine variously assigned to Versoporcus steinheimensis or ‘Parachleuastochoerus’ valentini. Our revision of the available remains, including unpublished material recovered from the 1980s onward, confirms the presence of the two former species and the assignment of the large tetraconodontine to ‘Pa.’ valentini. On biochronological grounds, the association of these taxa is characteristic of the early Vallesian, with the record of L. splendens representing the last well-dated occurrence of this species. From a paleoenvironmental viewpoint, the suid assemblage from CLL1 is consistent with previous inferences indicating the presence of a densely forested and humid environment with more open habitats nearby. Sara G. Arranz et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Sara G. Arranz Souzanna Siarabi Sharrah McKenzie Isaac Casanovas-Vilar David M. Alba Dental morphology Europe evolution Miocene Spain suids systematics taxonomy 2026-02-23T16:40:37+02:00 2026-02-23T16:40:40+02:00 2026-02-23T16:40:54+02:00 Adobe InDesign 21.1 (Windows) uuid:7474acad-c4f3-42e3-bd6f-35ebdd5eefb7 xmp.did:9F28E084BD8CE31184EC999A5D291C94 xmp.id:24d6fe6a-ab3a-e244-a5b5-6f85d242530e proof:pdf xmp.iid:809534ef-1ff7-174c-95ad-1de59944586f xmp.did:4444FC51923DE811ADBCA6C78E600139 xmp.did:9F28E084BD8CE31184EC999A5D291C94 default converted from application/x-indesign to application/pdf Adobe InDesign 21.1 (Windows) / 2026-02-23T16:40:37+02:00 Adobe PDF Library 18.0 False Pages: Page: Label: 267 Sequence: 1 Annotations: Annotation: Subtype: Link Rect: 51, 594, 268, 583 Annotation: Subtype: Link Rect: 180, 576, 243, 565 Annotation: Subtype: Link Rect: 303, 576, 362, 565 Annotation: Subtype: Link Rect: 302, 57, 435, 49 Page: Label: 268 Sequence: 2 Page: Label: 269 Sequence: 3 Page: Label: 270 Sequence: 4 Page: Label: 271 Sequence: 5 Page: Label: 272 Sequence: 6 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