Traits measured were head width (the widest part of the head including the compound eyes, HW); forewing: length of the forewing subcostal vein (SC), length of vannal vein, length of first medial cell (MEDIAL1), first cubital cell length, second submarginal cell length (MARG2); hind wing: length, cubital vein length, total wing length; and hind leg: basitarsus length, femur length (FEMUR), width of femoral-trochanter ridge.