Pollen cones grouped, ovate or oblong-cylindric, leaving gall-like protuberances after falling, yellow to red, green, blue, or purple. Seed cones maturing in 1 season, erect, ovoid to oblong-cylindric or cylindric, not falling whole but scale by scale, cone axis persisting as an erect "spike" on branch; scales shed individually, fan-shaped, lacking apophysis and umbo; bracts included to exserted.